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Morozovella acuta Toulmin 1941 from: Van Eijden, A.J.M.Smit, J. (1991): Eastern Indian Ocean Cretaceous and Paleogene quantitative biostratigraphy. In: Proceedings of the Ocean Drilling Program, Scientific Results Vol. 121 Eds: Weissel, J.Peirce, J.Taylor, E.Alt, J. p. 77-123
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Species Morozovella acuta Toulmin 1941



Discussion / Comments:
Toumarkine & Luterbacher (1985):
The conicotruncate test of this species has a well developed 'keel' and ornamented umbilical shoulders at least in the early chambers of the last whorl. The last chamber occupies up to one-third of the whorl.
Pearson et al. (2006):
DISCUSSION.- This taxon was discussed in the Atlas of Paleocene Planktonic Foraminifera (Olsson and others, 1999). STRATIGRAPHIC RANGE.- Zone P4b to Zone E2 (top).
Systematics:

1
 Superregnum Eukaryota
  Regnum Protoctista
   Phylum Ciliophora
    Subphylum Postciliodesmatophora
     Ordo Globigerinida
      Superfamilia Globorotaliaceae
       Superfamilia Nonionacea
        Familia Truncorotaloididae
         Genus Morozovella
          Species Morozovella acuta

15
  Classis Foraminifera
   Genus Morozovella
    Species Morozovella acuta

17
 Classis Foraminifera
  Genus Morozovella
   Species Morozovella acuta

19
  Ordo Foraminiferida
   Superfamilia Globorotaliaceae
    Familia Truncorotaloididae
     Genus Morozovella
      Species Morozovella acuta

32
 Ordo Foraminiferida
  Familia Globorotaliidae
   Genus Morozovella
    Species Morozovella acuta

35
  Ordo Foraminiferida
   Superfamilia Globigerinaceae
    Familia Truncorotaloididae
     Genus Morozovella
      Species Morozovella acuta
Synonym list:
Berggren & Norris (1997):
1941 Globorotalia wilcoxensis var. acuta Toulmin. - Toulmin : p. 608 pl. 82, figs. 6-8 (Zone P4, Salt Mountain Limestone Formation. Alabama).
1942 Globorotalia wilcoxensis var. acuta Toulmin. - Cushman & Renz : p. 12 pl. 3, figs. 2a-c (Zone P4, Soldado Formation, Trinidad).
1944 Globorotalia wilcoxensis var. acuta Toulmin. - Cushman : p. 48, 49 pl. 8, figs. 5a-b (Zone P3b, Naheola Formation, Alabama).
1944 Globorotalia wilcoxensis var. acuta Toulmin. - Cushman : p. 15 pl. 2, figs. 16a-b (Zone P5, Bashi Formation, Wilcox Group, Alabama).
1948 Globorotalia wilcoxensis var. acuta Toulmin. - Shifflett : p. 73 pl.4, figs. 23a-c (Zone P4, Aquia Formation, Wilcox Group, Maryland).
1954 Globorotalia velascoensis var. parva Rey. - Rey : p. 209 pl. 12, figs. la,b (Zone P4, Sample TB 450, Well NO. 11, Koudiat Bou-Khelif, near Ouezzane, N Morocco).
1957 Globorotalia acuta Toulmin. - Loeblich & Tappan : p. 185, 186 pl. 47, figs. 5a-c (Zone P4, Salt Mountain Limestone Formation, Wilcox Group, Alabama); pl. 55, figs. 4a-5c (Zone P4, Vincentown Formation, New Jersey); pl. 58, figs. 5a-c (Zone P4, Aquia Formation, Virginia).
1957 Globorotalia rex Martin. - Loeblich & Tappan (non Martin, 1943) : p. 195 pl. 60, figs, la-c (Zone P4, Ostrea thirsae Beds, Nanafalia Formation, Wilcox Counbty, Alabama).
1960 Globorotalia acuta Toulmin. - Berggren : pl. 1, figs. 15a-c (biostratigraphically equivalent to Zone P4, Ilaro, Nigeria).
1960 Globorotalia acuta Toulmin. - Reyment : p. 79-80 pl. 15, figs. 12a-b, 13; pl. 16, fig. 9 (upper Paleocene, W Nigeria).
1960 Globorotalia velascoensis parva Rey. - Bolli & Cita : p. 392-393 pl. 35, figs. 5A-C (Zone P4, Paderno d"Adda, N Italy).
1961 Globorotalia (Truncorotalia) acuta Toulmin. - Küpper : p. 257-258 pl. 16, figs. 3a-c (Zone P4, Zistersdorf. Austria, lower Ilerdian Stage).
1962 Globorotalia velascoensis parva Rey. - Gartner & Hay : p. 565-566 pl. 2, figs. 5a-c (Zone P4, Tremp Basin, Lerida, Spain, type Ilerdian Stage).
1963 Truncorotalia velascoensis acuta Toulmin. - Gohrbandt : p. 61 pl. 4, figs. 13-15 (Zone E=Zone P4, Salzburg Basin, Austria).
1963 Truncorotalia velascoensis parva Rey. - Gohrbandt : p. 61,62 pl. 5, figs. 4-6 (Zone E=Zone P4, Salzburg Basin, Austria).
1964 Globorotalia acuta Toulmin. - Luterbacher : p. 686-689 textfigs. 10la-c (Zone P4, El Quss Abu Said, Farafrah Oasis, Egypt); text-figs. 102a-104-c (Zone P5, Velasco Formation. Ebano, E Mexico).
1964 Globorotalia velascoensis parva Rey. - Luterbacher : p. 678 text-figs.9la-c ((re)illustration of supposed holotype).
1965 Globorotalia velascoensis forma acuta Toulmin. - Proto Decima & Zorzi : p. 34 pl. 4, figs. 23a-c; pl. 5, fig. 6 (Zone P4, Molinetta di Pederobba section, W Trevigiano, Italy).
1966 Globorotalia acuta Toulmin. - El Naggar : p. 188-190 pl. 19, figs. 5a-c ; pl. 20, figs, la-c (Zone P4, Lower Owaina Shale, Gebel Owaina, Egypt).
1966 Globorotalia velascoensis parva Rey. - El Naggar : p. 244,245 pl. 20, figs. la,b (Zone P4, Lower Oweina Marl, Esna Group, Gebel Oweina, Egypt).
1970 Globorotalia acuta Toulmin. - Samanta : p. 615-617 pl. 97, figs. 1,2 (Zone P4, Pondicherry Formation, upper Marlstone Unit, S India).
1970 Globorotalia velascoensis acuta Toulmin. - Shutskaya : p. 119-120 pl. 27, figs, lla-c (Ac. acarinata Tone., Tarkhankut Peninsula, Crimea, Kachan Stage); pl. 28, figs. 4a-c and pl. 29, figs. 9a-c (G. aequa Zone, Tarkhankut Peninsula, Crimea, Bakhchisarayan Stage).
1970 Globorotalia parva Rey. - Samanta : p. 628,629 pl. 97, figs. 3.4 (Zone P4, upper Marlstone Unit, Pondicherry Formation, Pondicherry area, S India).
1971 Globorotalia (Morozovella) acuta Toulmin. - Jenkins : p. 106 pl. 9, figs. 205-207 (G. triloculinoides Zone= Zone P4, Middle Waipara River section, Waipawan Stage, New Zealand).
1971 Globorotalia (Morozovella) velascoensis parva Rey. - Jenkins : p. 106, 107 pl. 9, figs. 211-213 (G. triloculinoides Zone=Zone P4, Middle Waipara River section, Waipawan Stage, New Zeland).
1975 Globorotalia acuta Toulmin. - Stainforth et al. : p. 163 text-figs. 30 (1-5) (Zone P4, Velasco Formation, Mexico); text-figs. 30 (6a-c) (Zone P4, Vincentown Formation, New Jersey=refigured illustrations of Loeblich and Tappan 1957, pl. 55, figs. 5a-c).
1977 Morozovella acuta Toulmin. - Berggren : p. 234 pl. 6 (reillustration of figured specimens in literature).
1977 Morozovella parva Rey. - Berggren : p. 236,237 pl. 7 (refigured illustrations from literature).
1979 Globorotalia (Morozovella) velascoensis parva Rey. - Blow : p. 1030, 1031 pl. 95, figs. 3-6 (Zone P5, Sample FRCM 1670, Lindi area, Tanzania, E Africa).
1985 Morozovella acuta Toulmin. - Toumarkine & Luterbacher : p. 114 text-figs. 14 (7; reillustration of holotype; 8: reillustration of LOEBLICH and TAPPAN 1957, pl. 55, figs. 4a-c, from the Vincentown Formation, New Jersey; incorrectly ascribed to the Salt Mountain Formation, Alabama=LOEBLICH and TAPPAN 1957, pl. 47, figs. 5a-c).
1997 Morozovella acuta Toulmin. - Berggren & Norris : p. 101 Plate 15 figures 19-21
Toumarkine & Luterbacher (1985):
1941 Globorotalia wilcoxensis var. acuta Toulmin. - Toulmin : p.608 pl 82 figs 6-8 (type reference)
1985 Morozovella acuta Toulmin. - Toumarkine & Luterbacher : p.111 figs 14.7-8
Olsson et al. (1999):
1941 Globorotalia wilcoxensis var. acuta Toulmin. - Toulmin : p. 608 pl. 82, figs. 6-8 (Zone P4, Salt Mountain Limestone Fm., Alabama)
1942 Globorotalia wilcoxensis var. acuta Toulmin. - Cushman & Renz : p. 12 pl. 3, figs. 2a-c (Zone P4, Soldado Fm., Trinidad)
1944 Globorotalia wilcoxensis var. acuta Toulmin. - Cushman : p. 48, 49 pl. 8, figs. 5a-b (Zone P3b, Naheola Fm., Alabama)
1944 Globorotalia wilcoxensis var. acuta Toulmin. - Cushman : p. 15 pl. 2, figs. 16a-b ( Zone P5, Bashi Fm., Wilcox Group, Alabama)
1948 Globorotalia wilcoxensis var. acuta Toulmin. - Shifflett : p. 73 pl.4, figs. 23a-c (Zone P4, Aquia Fm., Wilcox Group, Maryland).
1954 Globorotalia velascoensis var. parva Rey. - Rey : p. 209 pl. 12, figs. 1a,b (Zone P4, Sample TB 450, Well NO. 11, Koudiat Bou-Khelif, near Ouezzane, N Morocco)
1957 Globorotalia acuta Toulmin. - Loeblich & Tappan : p. 185, 186 pl. 47, figs. 5a-c (Zone P4, Salt Mountain Limestone Fm., Wilcox Group, Alabama); pl. 55, figs. 4a-5c (Zone P4, Vincentown Fm., New Jersey); pl. 58, figs. 5a-c (Zone P4, Aquia Fm., Virginia)
1960 Globorotalia velascoensis parva Rey. - Bolli & Cita : p. 392-393 pl. 35, figs. 5a-c (Zone P4, Paderno díAdda, N Italy).
1964 Globorotalia acuta Toulmin. - Luterbacher : p. 686-689 text- figs. 101a-c (Zone P4, El Quss Abu Said, Farafrah Oasis, Egypt); text-figs. 102a-104-c (Zone P5, Velasco Fm. Ebano, E Mexico)
1970 Globortalia velascoensis acuta Toulmin. - Shutskaya : p. 119-120 pl. 27, figs. 11a-c (Ac. acarinata Zone, Tarkhankut Peninsula, Crimea, Kachan Stage); pl. 28, figs. 4a-c and pl. 29, figs. 9a-c (G. aequa Zone, Tarkhankut Peninsula, Crimea, Bakhchisarayan Stage).
1971 Globorotalia (Morozovella) acuta Toulmin. - Jenkins : p. 106 pl. 9, figs. 205-207 ( G. triloculinoides Zone= Zone P4, Middle Waipara River section, Waipawan D Stage, New Zealand)
1971 Globorotalia (Morozovella) velascoensis parva Rey. - Jenkins : p. 106, 107 pl. 9, figs. 211-213 (G. triloculinoides Zone=Zone P4, middle Waipara River section, Waipawan Stage, New Zealand)
1979 Globorotalia (Morozovella) velascoensis parva Rey. - Blow : p. 1030, 1031 pl. 95, figs. 3-6 ( Zone P5, Sample FRCM 1670,Lindi area, Tanzania, E Africa).
1985 Morozovella acuta Toulmin. - Toumarkine & Luterbacher : p. 111 text-fig. 14 (7; reillustration of holotype; 8: reillustration of Loeblich and Tappan, 1957a, pl. 55, figs. 4a-c, from the Vincentown Fm., New Jersey; incorrectly ascribed to the Salt Mountain Fm., Alabama = Loeblich and Tappan, 1957a, pl. 47, figs. 5a-c).
Van Eijden & Smit (1991):
1941 Globorotalia wilcoxensis var. acuta Toulmin. - Toulmin : p.608 pl. 82, figs. 6-8
1991 Morozovella acuta Toulmin. - Van Eijden & Smit : p.113
Stratigraphy - absolute ages:
LAD: 54.7 ± 0 [Ma], Berggren et al. (1995) ODP Leg 80 Site 550
References:

Toulmin,L.D. (1941):
Eocene smaller foraminifera from the Salt Mountain limestone of Alabama . Journal of Paleontology Vol. 15 p. 567-611

Cushman,J.A. and Renz,H.H. (1942):
Eocene, Midway, Foraminifera from Soldado Rock, Trinidad . Contributions from the Cushman Foundation for Foraminiferal Research Vol. 18 p. 1-20

Cushman,J.A. (1944):
Foraminifera from the Aquia Formation of Virginia . Contributions from the Cushman Foundation for Foraminiferal Research Vol. 20 p. 17-28

Cushman,J.A. (1944):
A Paleocene Foraminiferal Fauna from the Coal Bluff Marl Member of the Naheola Formation of Alabama . Contributions from the Cushman Foundation for Foraminiferal Research Vol. 20 p. 29-49

Cushman,J.A. (1944):
A Foraminiferal Fauna of the Wilcox Eocene, Bashi Formation, from near Yellow Bluff, Alabama . American Journal of Sciences Vol. 242 p. 7-18

Shifflett,E. (1948):
Eocene Stratigraphy and Foraminifera of the Aquia Formation. Maryland . Department Geology, Mines and Water Resources Bulletin Vol. 3 p. 1-93

Rey,M. (1954):
Description de quelques espèces nouvelles de Foraminifères dans le Nummulitique nord-marocain . Bulletin Société Géologique de France Vol. 6(4) p. 209-211

Loeblich,A.R. and Tappan,H. (1957):
Planktonic Foraminifera of Paleocene and Early Eocene Age from the Gulf and Atlantic Coastral Plains.
In: Studies in Foraminifera, United States National Museum Bulletin Vol. 215 p. 173-198

Berggren,W.A. (1960):
Some planktonic Foraminifera from the Lower Eocene (Ypresian) of Denmark and northwestern Germany . Stockholm Contribution in Geology Vol. 5 p. 41-108

Reyment,R.A. (1960):
Notes on some Globigerinidae, Globotruncanidae and Globorotaliidae from the upper Cretaceous and lower Tertiary of western Nigeria . Records of the Geological Survey of Nigeria p. 68-86

Bolli,H.M. and Cita,M.B. (1960):
Globigerine e Globorotalie del Paleoceno di Paderno d'Ada (Italia) . Rivista Italiana Paleontologia Vol. 66(3)

Küpper,I. (1961):
Alttertiäre Foraminiferenfaunen in Flyschgesteinen aus dem Untergrund des nördlichen Inneralpinen Wiener Beckens (Österreich). , Jahrbuch der Geologischen Bundesanstalt Vol. 104

Gartner,S. and Hay,W.W. (1962):
Planktonic Foraminifera from the type Llerdian . Ecoglae geologicae Helvetiae Vol. 55 p. 553-572

Gohrbandt,K.H.A. (1963):
Zur Gliederung der Paläogen im Helvetikum nördlich Salzburg nach planktonischen Foraminiferen. I. Teil: Paleozän und tiefstes Untereozän . Mitteilungen der Geologischen Gesellschaft in Wien Vol. 56 p. 1-116

Luterbacher,H.P. (1964):
Studies in some Globorotalia from the Paleocene and Lower Eocene of the Central Apennines . Ecoglae geologicae Helvetiae Vol. 57 p. 631-730

Proto Decima,F. and Zorzi,P. (1965):
Studio micropaleontologico-stratigrafico della serie Cretaceo-Terziaria del Molinetto di Pederroba (Treviggiano occidentale) . Memorie degli Instituti di Geologia e Mineralogia dell'Universita di Padova Vol. 25 p. 44pp

El Naggar,Z.R.M. (1966):
Stratigraphy And Planktonic Foraminifera of the Upper Cretaceous - Lower Tertiary Succession in the Esna-Idfu Region. Nile Valley, Egypt . Egypt. Bull. Geol.Supp 2 p. 279 pp

Shutskaya,E.K. (1970):
Morfologicheskie grupirovki vidov rodov Globigerina i Acarinina v nizhnei chasti paleogena Kryma, Predkavkazya i zapada Srednei Azii i ipisanie vidov.
In: Stratigrafiya i paleontologiya Mezozoiskikh i Paleogenovykh otlozhenii Srednei Azii. Vsesoyuznyi Naucho-Issledovatel'skii Geologorazvedochnyi Nefyanoi Institut (VNIGNI), Trudy Vol. 69 Eds: Shutskaya, E.K. p. 79-134

Samanta,B.K. (1970):
Middle Eocene Planktonic Foraminifera from Lakhpat, Cutch, Western India . Micropaleontology Vol. 16

Jenkins,D.G. (1971):
New Zealand Cenozoic Planktonic Foraminifera . New Zealand Geological Survey Paleontological Bulletin Vol. 42

Stainforth,R.M.; Lamb,J.L.; Luterbacher,H.P.; Beard,J.H. and Jeffords,R.M. (1975):
Cenozoic planktonic foraminiferal zonation and characteristics of index forms . Paleontological ContributionsArticle 62 p. 425

Berggren,W.A. (1977):
Atlas of Paleogene planktonic foraminifera. Some species of the genera Subbotina, Planorotalites, Morozovella, Acarinina and Truncorotaloides.
In: Oceanic Micropaleontology Eds: Ramsay, A.T.S. p. 205-299

Blow,W.H. (1979):
The Cainozoic Globigerinida. 3 Vols p. 1413 pp

Toumarkine,M. and Luterbacher,H.P. (1985):
Paleocene and Eocene Planktic Foraminifera.
In: Plankton Stratigraphy p. 87-154

Van Eijden,A.J.M. and Smit,J. (1991):
Eastern Indian Ocean Cretaceous and Paleogene quantitative biostratigraphy.
In: Proceedings of the Ocean Drilling Program, Scientific Results Vol. 121 Eds: Weissel, J.Peirce, J.Taylor, E.Alt, J. p. 77-123

Berggren,W.A.; Kent,D.V.; Swisher,C.C. and Aubry,M.P. (1995):
A revised Cenozoic Geochronology and Chronostratigraphy.
In: Geochronology Time Scales and Global Stratigraphic Correlation, SEPM Special Publication Vol. 54

Berggren,W.A. and Norris,R.D. (1997):
Biostratigraphy, phylogeny and systematics of Paleocene trochospiral planktic foraminifera . Micropaleontology Vol. 43(suppl. 1) p. 1-116

Olsson,R.K.; Berggren,W.A.; Hemleben,C. and Huber,B.T. (1999):
Atlas of Paleocene planktonic foraminifera - online version. , Smithsonian Contributions to Paleobiology Vol. 85 p. 1-252

Pearson,P.N.; Olsson,R.K.; Hemleben,C.; Huber,B.T. and Berggren,W.A. (2006):
Atlas of Eocene Planktonic Foraminifera. p. 1-513

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