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Hedbergella infracretacea Glaessner 1937 from: Leckie, M.R. (1990): Middle Cretaceous Planktonic Foraminifers of the Antarctic Margin: Hole 693A, ODP Leg 113. In: Proceedings of the Ocean Drilling Programme Vol. 113 Eds: Parker, P.F..Kennett, J.P.O'Connell, S..Pisias, N.G. p. 319-324
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Species Hedbergella infracretacea Glaessner 1937



Alternative name:
Diagnosis / Definition:
Leckie (1990):
Test small, maximum diameter to 0.31 mm. Five to six subspherical to ovoid chambers in the final whorl, increasing gradually in size as added. The dorsal side varies from a low trochospire to flat; low trochospire tends to be more prevalent in juveniles and five-chambered adults, although the populations display a continuous range of variation. Test surface perforate and slightly pustulose. Pores occur at the apices of the pustules. Pustules on earlier chambers of final whorl occur in aggregates yielding subtle patterns suggestive of rugae. Aperture extraumbilical-umbilical; developed as a moderate arch with a thin imperforate rim
Discussion / Comments:
Leckie (1990):
The subtle "rugosity" developed in this taxon differs distinctly from the pattern in Hedbergella libyca or species of Rugoglobiqerina; the rugae of these latter taxa are non-perforate, the pores occur between rugae rather than being developed upon the ornament. This particular surface texture has been observed on several hedbergellids of late Aptian to early Albian age (e.g., H. trocoidea and H. delrioensis of Krasheninnikov and Basov, 1983; H. delrioensis of Longoria, 1974; Hedbergella sp. 1, H. aff. trocoidea, H. gorbachikae and H. cf. rischi of Leckie, 1984; Blefuscuiana cf. aptica of Banner and Desai, 1988). It is possible that the "rugosity" is a preservational artifact of hedbergellids of this age, although Leckie (in press) has argued against widespread poor preservation as the principal cause for the marked decline in planktonic foraminiferal diversity during latest Aptian-early Albian time. Another possibility is that it represents primary ultrastructure and therefore is of Phylogenetic significance. Hedbergella infracretacea (Glaessner) has been a problematic taxon, especially with regard to its relationship with H. delrioensis (Carsey) (see discussions by Hermes, 1969; Longoria, 1974; Masters, 1977; Price, 1977). Glaessner (1966) attempted to clarify the species concept of H. infracretacea by illustrating three topotypes, all of which clearly show the absence of an apertural lip. This observation is ultimately what led the present author to assign the Hole 693A specimens to the H. infracretacea group. Glaessner (1937) also notes that this taxon has five to six chambers in its final whorl. Other diagnostic characteristics emphasized by other authors include the small size of H. infracretacea and its variable dorsal convexity. H. delrioensis s.s. is characterized by having four and one-half to five and one-half chambers in its final whorl, a flat dorsal side, a hispid test, and in possessing a spatulate lip. The populations from Hole 693A show a continuous range of variability in terms of size (up to 0.31 mm), dorsal convexity (moderately elevated to depressed early coil), and number of chambers in the final whorl (5-6). However, no specimens were observed with a "hispid" test or with an apertural lip, and therefore classification within H. delrioensis was rejected. H. infracretacea and H. delrioensis probably co-occur in some sediments of late Aptian to middle Albian age, but H. delrioensis s.s. is best developed in late Albian and Cenomanian assemblages (e.g., Price, 1977; Leckie, 1984). Hedbergella infracretacea differs from H planispira (Tappan) in lacking a well developed apertural lip and in lacking the characteristic circular outline of that taxon. In addition, H. planispira has up to eight chambers in the final whorl. Specimens of H. infracretacea from Hole 693A may include forms referable to H. rischi Moullade (H. aff. infracretacea of Moullade, 1966).
Pflaumann & Krasheninnikov (1977):
H. infracretacea contains five to five and one-half ovoidal chambers in the last whorl which increases very slightly, umbilicus deep and narrow. In Hole 369A samples five-chambered specimens are dominant in the late Aptian section, while in the Albian five and one-half chambers are more common. Differences against H. planispira: higher trochospiral, and narrow umbilicus, against H. trocoidea: lower trochospiral and lower chamber number, smoother surface, against H. delrionensis: last chamber shows no protrusion towards the umbilicus, ovoid in shape with longest axis in direction of growth.
Synonym list:
Leckie (1990):
1937 Globigerina infracretacea Glaessner. - Glaessner : p. 28 text-fig. 1.
1966 Hedbergella infracretacea Glaessner. - Glaessner : p. 181-185 pl.1, figs. la-3c
p 1984 Hedbergella delrioensis Carsey. - Leckie : p. 598 pl.1, fig. 12, non: pl, 9, figs. 1-4, 8 early forms
1990 Hedbergella infracretacea Glaessner. - Leckie : Pl. 2, Figs. 1-18 as Hedbergella infracretacea group
Pflaumann & Krasheninnikov (1977):
1937 Globigerina infracretacea Glaessner. - Glaessner : p.28 text-fig. 1
1966 Hedbergella infracretacea Glaessner. - Glaessner : p.179 pl. 1; fig. 1-3
Was used in synonym list of:
References:

Glaessner,M.F. (1937):
Planktonische Foraminiferen aus der Kreide und dem Eozän und ihre stratigraphische Bedeutung.
In: Studies in Micropaleontology Vol. 1(1) p. 27-46

Glaessner,M.F. (1966):
Notes on foraminifera of the genus Hedbergella . Ecoglae geologicae Helvetiae Vol. 59 p. 179-185

Pflaumann,U. and Krasheninnikov,V.. (1977):
Early Cretaceous planktonic foraminifers from eastern North Atlantic, DSDP Leg 41 .
In: Initial Reports of the Deep Sea Drilling Project 41 , Abidjan, Ivory Coast, to Malaga, Spain, February-April 1975 Eds: Gardner, J..Herring, J.. p. 539-564

Leckie,M.R. (1984):
Mid-Cretaceous planktonic foraminiferal biostratigraphy off Morocco, Deep Sea Drilling Project Leg 79, Sites 545 and 547 . Initial Reports of the Deep Sea Drilling Project Vol. 79 p. 579–620

Leckie,M.R. (1990):
Middle Cretaceous Planktonic Foraminifers of the Antarctic Margin: Hole 693A, ODP Leg 113.
In: Proceedings of the Ocean Drilling Programme Vol. 113 Eds: Parker, P.F..Kennett, J.P.O'Connell, S..Pisias, N.G. p. 319-324

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