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Struthiolaria ornata Sowerby 1846 from: Griffin, M..Nielsen, S.N.. (2008): A revision of the type specimens of Tertiary molluscs from Chile and Argentina described by d’Orbigny (1842), Sowerby (1846), and Hupé (1854) . Journal of Systematic Palaeontology
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Species Struthiolaria ornata Sowerby 1846



Discussion / Comments:
Griffin & Nielsen (2008):
TYPE MATERIAL. Syntype: NHM-G26392 (Santa Cruz). REMARKS. It appears that Sowerby (1846) had a variety of specimens from different localities, but the figured specimen seems to be from Santa Cruz, as suggested by the label in the NHM; the specimen, however, is missing from the Darwin Collection. This is in contradiction to Camacho & Zinsmeister’s (1989, p. 105) statement that the “holotype” was an internal mold from the San Julián Formation at Cabo Curioso. No indication is given as to how they reached such a conclusion, but statements in the original publication by Sowerby leave no doubts about the existence of syntypes, rendering such a “designation of lectotype by inference of holotype” invalid (ICZN 1999, Art. 74.5). In fact, Sowerby himself stated in the original description that “Casts of a large variety are found in a loose clayey sandstone at Port San Julian”, a statement that clearly indicates that the species (i.e., the “typical” specimens) came from the Monte León Formation, which is also exposed around Port San Julián. However, a similar species also occurs in the San Julián Formation. Thus, the main complication that clouds the identity of this taxon is the fact that the species from Santa Cruz (probably, although impossible to confirm, as the material is missing in the Darwin Collection) – occurring there and at other localities including San Julián but always within the Monte León Formation – is different from the one common in the San Julián Formation. A lectotype designation would be necessary in order to unequivocally identify them both. Unfortunately, the specimen in the NHM is a very poorly preserved internal mould, which is very difficult to assign to either of the two species. Thus, an application to the ICZN (Art. 75.5) would be necessary in order to set that specimen aside and designate a neotype from the Monte León Formation, of which numerous well preserved specimens are available. Cenozoic South American struthiolariids have been generally placed in the genus Struthiolarella Steinmann & Wilckens, 1908 [type species: Struthiolaria ameghinoi Ihering, 1897]. However, we follow Powell (1951) and Nielsen (2005) in regarding Struthiolarella as a synonym of Perissodonta Martens, 1878 [type species Struthiolaria mirabilis Smith, 1875].
Synonym list:
Griffin & Nielsen (2008):
1846 Struthiolaria ornata Sowerby. - Sowerby : p.250 pl. 4; fig. 62
2008 Perissodonta ornata Sowerby. - Griffin & Nielsen : pl. 22; fig. 9
References:

Sowerby,G.B.. (1846):
Descriptions of Tertiary fossil shells from South America..
In: Geological observations on South America. Eds: Darwin, C.. p. 249-264

Griffin,M.. and Nielsen,S.N.. (2008):
A revision of the type specimens of Tertiary molluscs from Chile and Argentina described by d’Orbigny (1842), Sowerby (1846), and Hupé (1854) . Journal of Systematic Palaeontology

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