Hantkenina australis Finlay 1939 from: Pearson, P.N.Olsson, R.K.Hemleben, C.Huber, B.T.Berggren, W.A. (2006): Atlas of Eocene Planktonic Foraminifera. p. 1-513 . |
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Species Hantkenina australis Finlay 1939 |
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Alternative name: |
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Diagnosis / Definition: |
Pearson et al. (2006):
DESCRIPTION.
Type of wall: Smooth, normal perforate,
probably nonspinose; average pore size is often smaller
than in other species of Hantkenina; tubulospines
imperforate, smooth or with spiral rifling.
Test morphology: Planispiral, laterally
compressed with 5-6 closely appressed subtriangular
chambers in the final whorl; peripheral outline lobed or
slightly angular, anterior chamber shoulder is minimal
or non-existent; most or all of the adult chambers extend
into a hollow tubulospine; primary aperture is an
equatorial high arch extending about halfway up the
apertural face, widening towards the base into weak
basal lobes and bordered on each side by an imperforate
lip; sutures depressed, straight or slightly sigmoidal;
pustules common on early chambers of the final whorl
and in the umbilical region; tubulospines slender, often
long, curved backwards slighily in the opposite direction
to coiling, tapering to a poiilt, arising sharply from the
supporting chamber, positioned at or just spanning the
anterior chamber suture, sometimes partially contacting
the adjacent younger chamber.
Size: Maximum diameter without spines: 0.45
mm, with spines, 0.56 mm (Jenkins, 1965). |
Discussion / Comments: |
Jenkins (1985):
H. australis is distinguished from other species of the genus by recurved spines. It is found sporadically in the Middle Late Eocene Globigerinatheka index Globorotalia inconspicua zones in New Zealand. The only other good record of H. australis is in the early Middle Eocene of DSDP Leg 40 Site 362A in the South Atlantic (Tournarkine, 1978).
Pearson et al. (2006):
DISTINGUISHING FEATURES.-
This species has a
variable test morphology, showing features of H.
dumblei and H. compressa. It differs froin both, and all
other species of Hantkenina in having posteriorly
recurved tubulospines.
DISCUSSION.-
Finlay (1939) did not provide a
detailed description or designate a type specimen when
he erected this species because the available specimens
were incomplete. He stated that the holotype would be
figured later when new material was found and instead
figured a paratype from a different locality. This
specimen differs significantly from the type description
in lacking recurved tubulospines and having a more
angular outline.
Jenkins (1965) produced the first detailed
description of H. australis and selected a type specimen
out of the six incomplete specimens from Finlay's
original sample that satisfactorily represents the taxon.
This specimen is illustrated in SEM for the first time
(Pl. 8.5, Figs. 1-2). Brönnimann (1950) and Ramsay
(1962) recorded H. australis in Trinidad and Tanzania
respectively but in both cases the identification was based on Finlay's paratype and the illustrated specimens
do not have recurved tubulospines.
Our concept of H. australis is based on Finlay's
(1939) and Jenkins's (1965) notion of a form with
backward curving tubulospines. Subbotina's (1953)
illustrations of 'Hantkenina alabamensis' clearly show
this distinctive feature. We have found this species in
correlatable sequences hom the southern Labrador Sea
(ODP Site 647), Uzbekistan, southern Russia and the
Ukraine (Beniamovski, pers. comrn., 2001), indicating
that it has a global distribution. It appears to be most
common at the high southerly and northerly extremes
of the hantkeninid latitudinal range suggesting it was
more tolerant of cold water than other hantkeninids. In
parallel with the H. dmmblei-H. compressa-H.
alahamensis transition, there is a tendency for the test
to become more inflated through time. Hantkenina
compressa coexists with H. australis in the upper middle
Eocene in New Zealand. A common feature of this taxon
is for tubulospines to be absent on the early chambers
as in H. primitiva, which makes it impossible to
distinguish between them when the tubulospines are
missing.
PHYLOGENETIC RELATIONSHIPS.-
Probably evolved from H. dumblei in the middle Eocene.
STRATIGRAPHIC RANGE.-
Middle Eocene, Zones Ell-E13.
GEOGRAPHIC DISTRIBUTION.-
Global, low to mid latitudes, most common at the high northerly and
southerly extremes of the hantkeninid range, i.e., New
Zealand, southern Labrador Sea.
STABLE ISOTOPE PALEOBIOLOGY.-
No data available. |
Systematics: |
35 Ordo Foraminiferida
Superfamilia Globigerinaceae
Familia Hantkeninidae
Genus Hantkenina
Species Hantkenina alabamensis
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Synonym list: |
Jenkins (1985):
Pearson et al. (2006):
1939 Hantkenina australis Finlay. - Finlay : p.538 pl. 56; fig. 20, 21 [middle Eocene, Hampden, New Zealand]
1953 Hantkenina alabamensis Cushman. - Subbotina : p.146 pl. 1; fig. 6-7 [upper Eocene, northern Caucasus,
Russia]; [Not Cushman, 1925]
1965 Hantkenina australis Finlay. - Jenkins : p.518 fig. 1, 2 [late middle Eocene, Hampden, New
Zealand, sample F5 179B
2006 Hantkenina australis Finlay. - Pearson et al. : p.231 pl. 8.5; fig. 1-15 (Pl. 8.5, Figs. 1-2: new SEMs of the holotype of
Hantkenina australis Finlay)
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Was used in synonym list of: |
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Specimen: |
New Zealand Geological Survey Collection - Lower Hutt, New Zealand, Inventory number: 103211
New Zealand Geological Survey Collection - Lower Hutt, New Zealand, Inventory number: 103213
New Zealand Geological Survey Collection - Lower Hutt, New Zealand, Inventory number: F5601
New Zealand Geological Survey Collection - Lower Hutt, New Zealand, Inventory number: 1035/4-6
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References: |
Finlay,H.J. (1939): New Zealand foraminifera; key species in stratigraphy, No. 3 . Transactions of the Royal Society of New Zealand Vol. 69 p. 309-29
Finlay,H.J. (1939): New Zealand foraminifera; key species in stratigraphy, No. 1 . Transactions of the Royal Society of New Zealand Vol. 68
Thalmann,H.E. (1942): Foraminiferal genus Hantkenina and its subgenera . American Journal of Sciences Vol. 240 p. 809-823
Brönnimann,P. (1950): The Genus Hantkenina Cushman in Trinidad and Barbados, B.W.I. . Journal of Paleontology Vol. 24 p. 397-420
Subbotina,N.N. (1953): Fossil foraminifera of the USSR. Globigerinidae, Hantkeninidae and Globorotaliidae [in Russian] . Trudy Vsesoyuznogo Neftyanogo Nauchno-Isledovatelskogo Geologo-Razvedochnogo Instituta (VNIGRI), Novaya Seriya Vol. 76 p. 1-296
Ramsay,W.R. (1962): Hantkenininae in the Tertiary rocks of Tanganyika . Contributions from the Cushman Foundation for Foraminiferal Research Vol. 13 p. 79-89
Jenkins,D.G. (1965): The genus Hantkenina in New Zealand . New Zealand Journal of Geology and Geophysics Vol. 8 p. 518-526
Jenkins,D.G. (1985): Southern mid-latitude Paleocene to Holocene planktic foraminifera. In: Plankton stratigraphy Eds: Bolli, H.M.Saunders, J.B.Perch-Nielsen, K. p. 263-282
Pearson,P.N.; Olsson,R.K.; Hemleben,C.; Huber,B.T. and Berggren,W.A. (2006): Atlas of Eocene Planktonic Foraminifera. p. 1-513
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