Clavigerinella columbiana Petters 1954 from: Pearson, P.N.Olsson, R.K.Hemleben, C.Huber, B.T.Berggren, W.A. (2006): Atlas of Eocene Planktonic Foraminifera. p. 1-513 . |
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Species Clavigerinella columbiana Petters 1954 |
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Diagnosis / Definition: |
Pearson et al. (2006):
DESCRIPTION.
Type of wall: Uncertain because all the
specimens we have examined are poorly preserved; it
may have a higher pore density than is typical of the
genus.
Test morphology: Planispiral, involute,
biumbilicate; 4-4 1/2 chambers in the final whorl,
increasing rapidly in size as added; early chambers
rounded, final 2-3 adult chambers radially elongate,
distinctly flattened in the direction of coiling into subtriangular
'paddles', arranged perpendicular to the
direction of coiling; individual chambers are wide at
the distal ends, tapering to a narrow waist at the point
of attachment; peripheral outline strongly lobulate; equatorial narrow arched aperture, symmetrical or slightly asymmetrical, bordered by an imperforate lip;
sutures straight or curved, short compared to the total length of chambers.
Size: Maximum diameter of holotype 0.66 mm;
minimum diameter 0.51 mm; maximum thickness (at
periphery) 0.40 mm; minimum thickness (near
umbilicus) 0.15 mm (Petters, 1954). Specimens reach
up to 1 mm in diameter. |
Discussion / Comments: |
Pearson et al. (2006):
DISTINGUISHING FEATURES.-
Clavigerinella
colombiana is differentiated from all other
clavigerinellids by having sub-triangular chambers that
are distinctly flattened in the anterior-posterior axis.
Petters likened the form of the test to a "paddlewheel".
DISCUSSION.-
Complete specimens are very rare;
the holotype illustrated by Petters, and shown here in
SEM for the first time, is one of very few that has been
isolated more or less intact. The species is usually
identified by the isolated, paddle-shaped chambers. A
few complete species have been reported from the
western equatorial Atlantic (Demerara Rise) (P. Sexton,
pers. comm., 2004).
PHYLOGENETIC RELATIONSHIPS.-
The
evolutionary relationships of C. colombiana to other
clavigerinellids is difficult to establish because of its
sporadic occurrence. It probably evolved from C.
eocanica in the latest early Eocene (E7) by lateral
widening and flattening of the chambers in the direction
of coiling.
STRATIGRAPIC RANGE.-
Uppermost Zone E7-E10 (poorly constrained).
GEOGRAPHIC DISTRIBUTION.-
Apparently
worldwide, most common in continental margin settings
and the equatorial Pacific. Not found in open-ocean
oligotrophic assemblages. It was originally described
from the same sample as Pseudoglobigerinella
bolivariana. Blow's (1 979) comment that this species
is particularly common in the Indo-Pacific region has
not been substantiated.
STABLE ISOTOPE PALEOBIOL0GY.-
Clavigerinella
colombiana registers high ò18O and low ò13C
compared to other co-occurring planktonic species,
indicating that it lived in a cold, 12C-rich water mass
(Coxall, 2000). |
Systematics: |
35 Ordo Foraminiferida
Superfamilia Globigerinaceae
Familia Hantkeninidae
Genus Clavigerinella
Species Clavigerinella columbiana
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Synonym list: |
Pearson et al. (2006):
1954 Hastigerinella colombiana Petters. - Petters : p.40 pl. 80; fig. 10a-b [middle Eocene, Asterigerina crassaformis zone, upper
Carreto Fm., Colombia]
1955 Hastigerinella colombiana Petters. - Weiss : p.309 pl. 2; fig. 9-10 [middle Eocene, Talara, Chira, and Mirador Fms,
Peru]
2006 Clavigerinella columbiana Petters. - Pearson et al. : p.220 pl. 8.2; fig. 9-13 (Pl. 8.2, Figs. 9-1 0: new SEMs of the holotype of
Hastigerinella colombiana Petters)
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Was used in synonym list of: |
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Specimen: |
Cushman Collection - Smithsonian Museum of Natural History, Washington, D.C., Inventory number: USNM 64577
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References: |
Petters,V. (1954): Tertiary and Upper Cretaceous foraminifera from Colombia, S.A. . Contributions from the Cushman Foundation for Foraminiferal Research Vol. 5 p. 37-41
Weiss,L. (1955): Planktonic index Foraminifera of northwestern Peru . Micropaleontology Vol. 1(4) p. 301-308
Pearson,P.N.; Olsson,R.K.; Hemleben,C.; Huber,B.T. and Berggren,W.A. (2006): Atlas of Eocene Planktonic Foraminifera. p. 1-513
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