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Subbotina jacksonensis Bandy 1949 from: Pearson, P.N.Olsson, R.K.Hemleben, C.Huber, B.T.Berggren, W.A. (2006): Atlas of Eocene Planktonic Foraminifera. p. 1-513
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Species Subbotina jacksonensis Bandy 1949



Diagnosis / Definition:
Pearson et al. (2006):
DESCHPTION. Type of wall: Normal perforate, cancellate, spinose, ruber/sacculifier-type wall texture. Test morphology: Test low trochospiral, globular, ovate in outline, chambers globular, much embracing; in spiral view 4 globular, much embracing chambers in ultimate whorl, increasing moderately in size, ultimate chamber varying in size, sutures very slightly depressed, straight; in umbilical view 4 globular, much embracing chambers, increasing moderately in size, ultimate chamber smaller in size than penultimate chamber, directed over and covering the umbilicus, mature wall texture may or may not be developed on ultimate chamber, sutures very slightly depressed, straight, umbilicus small but hidden by ultimate chamber, aperture umbilical; in edge view low trochospire, chambers globular in shape, much embracing, ultimate chamber projected over the umbilicus. Size: Maximum diameter of holotype 0.39 mm, thickness 0.29 mm.
Discussion / Comments:
Pearson et al. (2006):
DISTINGUISHING FEATURES.- The species is characterized by its globular test, much embracing chambers, with a reduced ultimate chamber projecting over and covering the umbilicus. DISCUSSION.- We illustrate in SEM for the first time the holotype of Subbotina jacksonensis (Pl. 6.13, Figs. 1-3). This species has been apparently overlooked by workers, or grouped with other subbotinid species, or placed in Catapsydrax. The ultimate chamber, even though it projects over and covers the umbilicus, cannot be regarded as a bulla. It represents the terminal stage in the ontogeny of the individual and shows varying degrees of wall texture development among individuals of the species, a common feature in planktonic foraminifera. The species Sfairly common in our upper Eocene samples. Globigerina ouachitaensis senilis Bandy was described from a sample in the Yazoo Clay close to the sample from which S. jacksonensis was described. Globigerina senilis was described as having a small ultimate chamber that restricts the opening to the umbilicus, which is observed within the normal range of morphologic variation in S. jacksonensis. SEM images of Globigerina (Eoglobigerina) bizkaiensis Orue-Etxebarria from the middle Eocene of Spain are identical with holotype and other images of S. jacksonensis. PHYLOGENETIC RELATIONSHIPS.- Subbotina jachonensis probably evolved from Subbotina eocaena by developing more embracing chambers and a reduced ultimate chamber that projects over the umbilicus. GEOGRAPHIC DISTRIBUTION.- Global in low- to mid-latitudes. STABLE ISOTOPIC PALEOBIOLOGY.- No data available.
Systematics:

35
 Ordo Foraminiferida
  Superfamilia Globigerinaceae
   Familia Globigerinidae
    Genus Parasubbotina
     Species Parasubbotina varianta
      Species Subbotina jacksonensis
Synonym list:
Pearson et al. (2006):
1949 Globigerina rotundata jacksonensis Bandy. - Bandy : p.121 pl. 23; fig. 6a-c [upper Eocene, Yazoo Clay, Little Stave Creek, Clarke County, Alabama]
1949 Globigerina ouachitaensis senilis Bandy. - Bandy : p.121 pl. 22; fig. 5a-c [upper Eocene, Yazoo Clay, Little Stave Creek, Clarke County, Alabama]
non 1962 Globigerina senilis Bandy. - Blow & Banner : p.95 pl. XI; fig. r, s, t, u
1985 Globigerina (Eoglobigerina) bizkaiensis Orue-Etxebarria. - Orue-Etxebarria : p.469 pl. 1; fig. 1-10 [middle Eocene Globigerinatheka subconglobata and Glohorotalia (Morozovella) lehneri Zones, Province of Bizkaia, Spain]
2006 Subbotina jacksonensis Bandy. - Pearson et al. : p.143 pl. 6.13; fig. 1-20 (Pl. 6.13, Figs. 1-3: new SEMs of holotype of Globigerina rotundata jacksonensis Bandy) (Pl. 6.13, Figs. 4,7, 8: new SEMs of holotype of Globigerina ouachitaensis senilis Bandy)
Specimen:
Paleontologic Laboratory, Indiana University, Indiana, Inventory number: 4907
References:

Bandy,O.L. (1949):
Eocene and Oligocene foraminifera from Little Stave Creek, Clarke County, Alabama . Bulletins of American paleontology Vol. 32 p. 1-211

Blow,W.H. and Banner,F.T. (1962):
The Mid-Tertiary (Upper Eocene to Aquitanian) Globigerinaceae.
In: Fundamentals of Mid‑Tertiary Stratigraphical Correlation Eds: Eames, F.E.Banner, F.T.Blow, W.H.Clarke, W.J. p. 61‑151

Orue-Etxebarria,X.. (1985):
Descripcion de dos neuvas species de foraminiferos planctonicos en el Eoceno costero de la Provincia de Bizkaia . Revista Espaiiola de Micropaleontologia Vol. 17 p. 467-477

Pearson,P.N.; Olsson,R.K.; Hemleben,C.; Huber,B.T. and Berggren,W.A. (2006):
Atlas of Eocene Planktonic Foraminifera. p. 1-513

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