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Subbotina yeguaensis Weinzierl & Applin 1929 from: Pearson, P.N.Olsson, R.K.Hemleben, C.Huber, B.T.Berggren, W.A. (2006): Atlas of Eocene Planktonic Foraminifera. p. 1-513
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Species Subbotina yeguaensis Weinzierl & Applin 1929


Diagnosis / Definition:
Pearson et al. (2006):
DESCRIPTION. Type of wall: Normal perforate, symmetrically cancellate, sacculifer-type wall texture, spinose. Test morphology: Test moderately elevated, trochospiral, globular, lobulate in outline, chambers globular; in spiral view 3 1/3 globular, slightly embracing chambers in ultimate whorl, increasing moderately in size, ultimate chamber may be equal to or smaller in size than penultimate chamber, sutures moderately depressed, straight; in umbilical view 3 1/3 globular, slightly embracing chambers, increasing moderately in size, sutures moderately depressed, straight, umbilicus small, enclosed by surrounding chambers, aperture umbilical, bordered by a somewhat broad lip that tapers both anteriorly and posteriorly, ultimate chamber may be equal to or smaller than the penultimate chamber; in edge view moderately elevated trochospire, chambers globular in shape, somewhate embracing, aperture visible as a low munded opening, bordered by a somewhat broad lip. Size: Maximum diameter of holotype 0.52 mm, thickness 0.40 mm.
Discussion / Comments:
Pearson et al. (2006):
DISTINGUISHING FEATURES.- The species is characterized by its lobulate test and moderately elevated trochospire, and the somewhat broad apertural lip that tapers both anteriorly and posteriorly. DISCUSSION.- Blow (1979) included Subbotina yeguaensis in his new genus Dentoglobigerina, believing that it was an intermediate species between D. galavisi (Bermudez) and D. pseudovenezuelana (Blow and Banner). This was apparently based on his concept of a "central type" for D. galavisi, which are forms from the middle Eocene. However, these forms do not fit the central morphology of D. galavisi which was described from the upper Eocene Jackson Formation, Mississippi, which spans from Zones E 15- 16 (see holotype P1. 13.1, Figs. 1-3, this volume and discussion under D. galavisi). Subbotinayeguaensis is spinose and lacks the triangular lip that characterizes the nonspinose genus Dentoglobigerina and, therefore, cannot be placed as an intermediate species in a Dentoglobigerina lineage. Subbotina yeguaensis was described from the middle Eocene subsurface of Louisiana where it is common. The holotype of Subbotina pseudoeocaena (Subbotina) which is described from the lowerlmiddle Eocene of the northern Caucasus has the morphologic characteristics of S. yeguaensis and is considered a junior synonym (see P1. 6.18, Figs. 13-15), as is Subbotina angiporoides lindiensis of Blow (1979) and possibly Globigerina pseudoeocaena Subbotina var. ellipsocamerata Khalilov (1956; viewed by WAB). PHYLOGENETIC RELATIONSHIPS.- Subbotina yeguaensis belongs to a group of subbotinids with symmetrical cancellate (sacculifer-type) wall texture. It apparently arose in the early Eocene, possibly from S. eocaena. GEOGRAPHIC DISTRIBUTION.- Global in low to high latitudes. STABLE ISOTOPE PALEOBIOL0GY.- Boersma and others (1987) record strongly positive ò18O for this species indicating a deep planktonic habitat.
Systematics:

35
 Ordo Foraminiferida
  Superfamilia Globigerinaceae
   Familia Globigerinidae
    Genus Subbotina
     Species Subbotina yeguaensis
Synonym list:
Pearson et al. (2006):
1929 Globigerina yeguaensis Weinzierl & Applin. - Weinzierl & Applin : p.409 pl. 43; fig. 1a-b [middle Eocene, Yegua Fm., subsurface, Louisiana]
p 1953 Globigerina pseudoeocaena var. pseudoeocaena Subbotina. - Subbotina : p.81 pl. 11; fig. M. (partim; not pl. 5: figs. 1 a-2c = Subbotina eocaena; not pl. 5: fig. 6a-c = Subbotina corpulenta); [lower-middle Eocene Zone of conical Globorotalia, Kuban River, northern Caucasus].
? 1956 Globigerina pseudoeocaena var. ellipsocamerata Khalilov. - Khalilov : p.241 pl. 3; fig. 1a-c [middle Eocene, Maly Caucasus]
p 1957 Globigerina yeguaensis Weinzierl & Applin. - Bolli : p.163 pl. 35; fig. 15a-c (partim; not pl. 35: fig. 14a-c); [middle Eocene Zone E14, Navet Fm., Trinidad]
p 1962 Globigerina yeguaensis Weinzierl & Applin. - Saito : p.216 pl. 32; fig. 1a-c [middle Eocene Zone E12, Haha-Jima, Bonin Islands, western Pacific Ocean]. [Not pl. 32, figs. 2,3.]
1962 Globigerina linaperta pseudoeocaena Subbotina. - Blow & Banner : p.87 pl. 11; fig. M [upper Eocene Cribrohantkenina danvillensis Zone, Lindi area, Tanzania]
1969 Globigerina yeguaensis Weinzierl & Applin. - Blow : p.319 pl. 3; fig. 12-14 (holotype reillustraited)
1970 Globigerina yeguaensis Weinzierl & Applin. - Samanta : p.192 pl. 1; fig. 15-17 [middle Eocene Zone E12, Lakhpat, Cutch, western India]
1979 Globigerina yeguaensis Weinzierl & Applin. - Blow : p.1313 pl. 3; fig. 12-14 (holotype reillustrated)
1979 Subbotina pseudoeocaena Subbotina. - Blow : p.1279 pl. 187; fig. 8, 9 [Zone El011 1, DSDP Site 19, South Atlantic Ocean]
1979 Subbotina angiporoides lindiensis Blow. - Blow : p.1253 pl. 244; fig. 8-9 [upper Eocene Zone E15/16, Lindi, Tanzania]
1983 Globigerina pseudoeocaena Subbotina. - Krasheninnikov & Basov : p.838 pl. 2; fig. 1-3 [middle Eocene DSDP Site 512, Maurice Ewing Bank, South Atlantic Ocean].
p 1984 Subbotina pseudoeocaena Subbotina. - Belford : p.14 pl. 23; fig. 16-18 (partim; not pl. 23: figs. 19-21);[middle Eocene, Papua, New Guinea]
1985 Globigerina yeguaensis Weinzierl & Applin. - Bolli & Saunders : p.180 fig. 13-23a,b (holotype reillustrated)
1991 Subbotina yeguaensis Weinzierl & Applin. - Huber : p.441 pl. 5; fig. 2 [middle Eocene Zone AE314, ODP Site 738, Kerguelen Plateau, Southern Indian Ocean]
1994 Subbotina yeguaensis Weinzierl & Applin. - Nishi & Chaproniere : p.260 pl. 5; fig. 13-15 [upper Eocene Zone E141152 ODP Hole 841B, Tonga Trench, Pacific Ocean]
2001 Subbotina yeguaensis Weinzierl & Applin. - Warraich & Ogasawara : p.49 fig. 14: 4, 8, 12 [Zone E10/11, Kirthar Fm., Sulaiman Range, Pakistan]
2006 Subbotina yeguaensis Weinzierl & Applin. - Pearson et al. : p.152 pl. 6.18; fig. 1-16 (Pl. 6.18, Fig. 1-3: new SEM's of holotype of Globigerina yeguaensis Weinzierl and Applin)
Specimen:
Smithsonian Museum of Natural History, Washington, D.C., Inventory number: USNM CC12210
References:

Weinzierl,L.L. and Applin,E.R. (1929):
The Claiborne Formation on the Coastal Domes . Journal of Paleontology Vol. 3(4) p. 384-410

Subbotina,N.N. (1953):
Iskopaemye foraminifery SSSR (Globigerinidy, Khantkenininidy i Globorotaliidy) . Trudy Vsesoyznogo Nauchno-Issledovatel'skogo Geologo-razvedochnogo Instituta (VNIGRI) Vol. 76 p. 296

Khalilov,D.M. (1956):
O pelagicheskoy faune foraminifer Paleogenovykh otlozheniy Azerbaydzhana . Trudy Instituta Geologii, Akademiya Nauk Azerbaydzhanskoy SSR Vol. 17

Bolli,H.M. (1957):
Planktonic Foraminifera from the Eocene Navet and San Fernando formations of Trinidad, B.W.I. . Bull. U.S. natl. Mus. Vol. 215 p. 155-172

Blow,W.H. and Banner,F.T. (1962):
The Mid-Tertiary (Upper Eocene to Aquitanian) Globigerinaceae.
In: Fundamentals of Mid‑Tertiary Stratigraphical Correlation Eds: Eames, F.E.Banner, F.T.Blow, W.H.Clarke, W.J. p. 61‑151

Saito,T. (1962):
Eocene planktonic foraminifera from Hahajima (Hillsborough Island) . Trans. Proc. Paleontol. Soc. Japan, news series Vol. 45 p. 209-225

Blow,W.H. (1969):
Late middle Eocene to Recent planktonic foraminiferal biostratigraphy.
In: Proceedings of the First International Conference on Planktonic Microfossils, Geneva 1967 Vol. 1 Eds: Bronnimann, P.Renz, H.H. p. 199-422

Samanta,B.K. (1970):
Middle Eocene Planktonic Foraminifera from Lakhpat, Cutch, Western India . Micropaleontology Vol. 16

Blow,W.H. (1979):
The Cainozoic Globigerinida. 3 Vols p. 1413 pp

Krasheninnikov,V.A. and Basov,I.A. (1983):
Cenozoic planktonic foraminifers of the Falkland Plateau and Argentine Basin, Deep Sea Drilling Projejt Leg 71.
In: Initial Reports of the Deep Sea Drilling Project Vol. 71 Eds: Ludwig, W.J.Krasheninninov, V.A. p. 821-845

Belford,D.J. (1984):
Tertiary Foraminifera and Age of Sediments, Ok Tedi-Wabag, Papua, New Guinea . BureauMineral Resources, Geology and Geophysics, Australia Bulletin Vol. 216

Bolli,H.M. and Saunders,J.B. (1985):
Oligocene to Holocene low latitude planktic foraminifers.
In: Plankton Stratigraphy Eds: Bolli, H.M.Saunders, J.B. p. 155-262

Huber,B.T. (1991):
Paleogene and early Neogene planktonic foraminifer biostratigraphy of Sites 738 and 744, Kerguelen Plateau (southern Indian Ocean).
In: Proceedings of the Ocean Drilling Program, Scientific Results Vol. 119 Eds: Leckie, R.M.Sigurdsson, H.Acton, G.D.Draper, G. p. 427-449

Nishi,H.. and Chaproniere,G.C.H. (1994):
Eocene-Oligocene subtropical planktonic foraminifers at Site 841.
In: Proceedings of the Ocean Drilling Program, Scientific Results Vol. 135 Eds: Hawkins, J..Parson, L..Allan, J..others p. 245-266

Warraich,M.Y.. and Ogasawara,K.. (2001):
Tethyan Paleocene-Eocene planktic foraminifera from the Rakhi Nala and Zinda Pir land sections of the Sulaiman Range, Pakistan . Science Reports of the Institute of Geoscience University of Tsukuba, section B Vol. 22 p. 1-59

Pearson,P.N.; Olsson,R.K.; Hemleben,C.; Huber,B.T. and Berggren,W.A. (2006):
Atlas of Eocene Planktonic Foraminifera. p. 1-513
Anonymous: Unedited TaxonConcept data
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