Clavigerinella eocanica Nuttall 1928 from: Pearson, P.N.Olsson, R.K.Hemleben, C.Huber, B.T.Berggren, W.A. (2006): Atlas of Eocene Planktonic Foraminifera. p. 1-513 . |
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Species Clavigerinella eocanica Nuttall 1928 |
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Diagnosis / Definition: |
Pearson et al. (2006):
DESCRIPTION.
Type of wall: Usually smooth, normal perforate,
sometimes weakly cancellate; possibly spinose.
Test morphology: Planispiral or pseudoplanispiral,
evolute, laterally compressed biumbilicate
or showing a subtly raised spiral side and very shallow
umbilicus; 4-4 1/2 rapidly enlarging chambers in the final
whorl; early chambers rounded, final 2-3 adult chambers
conspicuously elongated in to club-shaped (clavate)
extensions; highly lobular peripheral outline; distal
chamber ends smoothly rounded; equatorial high arched
aperture, symmetrical or slightly asymmetrical, bordered
by a smooth broad imperforate lip, relict apertural lips
often present along sutures; sutures are shallow, straight,
becoming curved in later stages, short relative to total
chamber length.
Size: Maximum diameter 0.30-80 mm. |
Discussion / Comments: |
Pearson et al. (2006):
DISTINGUSHING FEATURES.-
Clavigerinella
eocanica is distinguished from all other clavigerinellids
in having unmodified, smoothly rounded clavate
chambers. It can be distinguished from its Miocene homeomorph Clavatorella bermudezi by the greater
lateral compression and shell symmetry (along the
equatorial plane), and fully equatorial position of the
aperture. It differs from Parasubbotina eoclava in
having better-developed clavate chambers, near
planispiral coiling, and a higher-arched, equatorially
positioned aperture. It differs from E! prebetica by the
lower, near-planispiral coiling, greater lateral
compression and possession of a high-arched, equatorial
aperture.
DISCUSSION.-
Toumarkine and Luterbacher (1985)
designated the specimen figured by Nuttall (1928) pl.
50: fig. 9 as the lectotype from three original illustrated
cotypes. On recent examination of Nuttall's
Hastigerinella eocanica (= Clavigerinella eocanica)
cotype suite in the USNM collection (11 in total),
however, no specimen could be found that exactly
matched this illustration, altdugh two are very similar
(i.e., same number, orientation and length of chambers).
Either the actual figured specimen is lost or the
illustration is inaccurate. The SEM image illustrated as
the probable lectotype of C. eocanica in this work (PI.
8.1, Figs. 10, 11) is the closest match among Nuttall's
cotypes to the illustration of the lectotype figured by
Toumarkine and Luterbacher (1985).
PHYLOGENTIC RELATIONSHIPS.-
This species
evolved from Parasubbotina eoclava in Zone E7 by
elongation of the chambers and lowering of the coil
height to form a planispiral. It diversified shortly after
its origin to produce a series of morphotypes including
C. caucasica, which was the ancestor of Hantkenina.
Detailed understanding of the relationship between C.
eocanica and other clavigerinellids is lacking because
of their sporadic geographic and stratigraphic
occurrence.
STRATIGRAPHIC RANGE.-
Upper Zone E7 to E 16.
This species is most characteristic of the lower middle
Eocene, but has been observed ranging into the upper
Eocene (Pearson and Chaisson, 1997).
GEOGRAPHIC DISTRIBUTION.-
Clavigerinella
eocanica is the most commonly encountered species of
the genus and has been found in open ocean as well as
marginal settings worldwide at mid-low latitudes.
STABLE ISOTOPE PALEOBIOL0GY.-
Clavigerinella
eocanica has high òl8O and low ò13C indicating
that it lived in a cold, deep possibly subthermocline
habitat (Pearson and others, 1993; Coxall and others,
2000). |
Systematics: |
35 Ordo Foraminiferida
Superfamilia Globigerinaceae
Familia Hantkeninidae
Genus Clavigerinella
Genus Hantkenina
Species Clavigerinella eocanica
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Synonym list: |
Pearson et al. (2006):
1957 Clavigerinella cf. akersi Bolli et al.. - Bolli : p.162 pl. 35; fig. 3a-b [Hantkenina aragonensis zone,
Navet Fm., Trinidad]; [Not Bolli, Loeblich and Tappan,
1957]
1971 Clavigerinella jarvisi Cushman. - Postuma : p.132 pl. on p. 133 (4 specimens, right hand column) [middle Eocene
Globorotalia bullbrooki Zone to Globigerapsis kugleri
Zone, Navet Fm., Trinidad]; [Not Cushman, 1930]
1976 Hastigerinella eocanica Nuttall. - Saito et al. : p.292 pl. 3; fig. 3a-c [middle Eocene, DSDP
Hole 2 1 A, South Atlantic Ocean]
1979 Clavigerinella eocanica eocanica Nuttall. - Blow : p. 1198-1203 pl. 151, fig. 1-3;
pl. 157, fig. 9;
pl. 162, fig. 1-3, 5-7 [Zone P9-P10, Kane 9-C Piston Core,
Endeavour Seamount, equatorial Atlantic Ocean]
1979 Clavigerinella cf. eocanica sensu lato Nuttall. - Blow : p. 1202-1203 pl. 151, fig. 4;
pl. 157, fig. 8 [Zone P9-
P10, Kane 9-C Piston Core, Endeavour Seamount,
equatorial Atlantic Ocean]
1985 Clavigerinella eocanica eocanica Nuttall. - Toumarkine & Luterbacher : p.119 pl. 22; fig. 2-3 [middle Eocene Morozovella lehneri Zone, El Datil Fm.,
Margarita, Venezuela].
1993 Clavigerinella eocanica Nuttall. - Pearson et al. : p.128 pl. 3; fig. 14, 16 [Middle Eocene Zone P1 1-
P12, DSDP Site 523, Walvis Ridge, South Atlantic
Ocean]
2003 Clavigerinella eocanica Nuttall. - Coxall et al. : p.238 pl. 1; fig. 9-14 [lower middle Eocene Zone P10, ODP Site 865, equatorial
Pacific Ocean]
2006 Clavigerinella eocanica Nuttall. - Pearson et al. : p.219 pl. 8.1; fig. 10-19 (Pl. 8.1, Figs. 10- 11 : new SEMs of the probable
lectotype of Hastigerinella eocanica Nuttall)
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Specimen: |
Smithsonian Museum of Natural History, Washington, D.C., Inventory number: USNM 517704
Smithsonian Museum of Natural History, Washington, D.C., Inventory number: USNM 517705
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References: |
Nuttall,W.L.F. (1928): Notes on the Tertiary foraminifera of Southern Mexico . Journal of Paleontology Vol. 2 p. 372-376
Bolli,H.M. (1957): Planktonic Foraminifera from the Eocene Navet and San Fernando formations of Trinidad, B.W.I. . Bull. U.S. natl. Mus. Vol. 215 p. 155-172
Postuma,J.A. (1971): Manual of Planktonic Foraminifera. p. 422 pp
Saito,T.; Thompson,P.R.. and Breger,D.. (1976): Skeletal ultramicrostructure of some clongate-chambered planktonic foraminifera and related species. In: Progress in Micropaleontology, sclected papers in honor of Professor Kiyoshi Asano Eds: Takayanagi, Y.Saito, T. p. 278-304
Blow,W.H. (1979): The Cainozoic Globigerinida. 3 Vols p. 1413 pp
Cremades Campos,J.. (1980): Eoclavatorella; nuevo genero de foraminifero planctonico del Eoceno inferior . Cuadernos de Geologia, Universidad de Granada Vol. 11 p. 209-214
Toumarkine,M. and Luterbacher,H.P. (1985): Paleocene and Eocene Planktic Foraminifera. In: Plankton Stratigraphy p. 87-154
Pearson,P.N.; Shackleton,N.J. and Hall,M.A. (1993): The Stable Isotope Paleoecology of Middle Eocene Planktonic Foraminifera and Multi-species integrated Isotope Stratigraphy . Journal of Foraminiferal Research Vol. 23 p. 123-140
Coxall,H.K..; Huber,B.T. and Pearson,P.N. (2003): Origin and morphology of the Eocene planktonic foraminifer Hantkenina . Journal of Foraminiferal Research Vol. 33 p. 237-261
Pearson,P.N.; Olsson,R.K.; Hemleben,C.; Huber,B.T. and Berggren,W.A. (2006): Atlas of Eocene Planktonic Foraminifera. p. 1-513
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