Hantkenina liebusi Shokhina 1937 from: Pearson, P.N.Olsson, R.K.Hemleben, C.Huber, B.T.Berggren, W.A. (2006): Atlas of Eocene Planktonic Foraminifera. p. 1-513 | . |
Notice: This catalogue page may contain unedited data.
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Species Hantkenina liebusi Shokhina 1937 |
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| Alternative name: |
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| Diagnosis / Definition: |
Pearson et al. (2006):
DESCRIPTION.
Type of wall: Smooth, normal perforate
probably nonspinose; tubulospines smooth or with spiral
rifling, imperforate or perforated by small sparsely
distributed pores with a well-defined zone of
demarcation between the tubulospines and the chamber wall.
Test morphology: Planispiral, involute,
biumbilicate and laterally compressed; 4-6 subtriangular chambers in the adult whorl, somewhat appressed, increasing rapidly in size as added; peripheral outline (excluding tubulospines) is slightly lobulate with minor
incisions between chambers; each chamber of the final
whorl extends into a hollow tubulospine; aperture is a
narrow, elongated equatorial arch bordered by an
imperforate lip that extends about two-thirds up the
apertural face, lip is often crenulated andlor pustulose
along its margin; sutures are depressed, straight,
becoming curved to slightly sigmoidal, remnants of
earlier apertural lips are sometimes present along the
sutures; tubulospines are constricted at the base, stout
on early adult chambers, longer and more slender in
final chambers, directed radially, positioned close to or
at the anterior chamber suture, long posterior chamber
shoulder and a minor or nonexistent anterior shoulder,
terminating in finger-like projections (coronet structure
of Ramsay, 1962) or tapering to a point, often with a
tiny circular terminal aperture.
Size: Maximum shell diameter (excluding
tubulospines) 0.48 mm (Shokhina, 1937). |
| Discussion / Comments: |
Pearson et al. (2006):
DISTINGUISHTNG FEATURES.-
Hantkenina liebusi
shares features of wall texture, aperture and tubulospines
with H. mexicana but differs in having a more
compressed test, more closely appressed chambers and
a less stellate peripheral outline. The tubulospines are
also in a more forward position with respect to the radial
chamber axis (i.e. there is a short anterior chamber shoulder).
Hantkenina liebusi usually has fewer
chambers in the final whorl than H. dumblei and H.
lekneri (4 1/2-6, compared to 5-7 in the latter). It can be
distinguished from H. dumblei by the slightly incised
peripheral outline and shorter adult chambers and from
H. lehneri in being generally less stellate.
DISCUSSION.-
The first Hantkenina to be illustrated was Hantken's 'Siderolina kochi'. This may be a prior
synonym for H. liebusi but the type specimen is lost and the original illustration is not definitive. When Shokhina (1937) described H. liebusi she specifically
rejected Hantken's figure as belonging to the same
species. Although Shokhina's specimens are also lost,
she illustrated the morphology quite clearly and it is
possible to make an almost exact match with specimens
from the middle Eocene (see Plate 8.9) which is the
level from which Shokhina described her species. The
holotype of H. longispina Cushman resembles
Shokhina's illustrations ofH. liebusi and may be a prior
synonym. It has not been possible to compare the type
specimens of these taxa because Shokhina's H. liebusi
material is unavailable for study and the type specimen
of H. longispina is missing from the USNM collections.
Moreover, H. longispina has been used indiscriminately
for a variety of only distantly related Hantkenina
morphotypes with 'long' tubulospines. Although the
name H. longispina cannot be rejected on the grounds
that it is an inappropriate name (ICZN, 1999, Art.
23.3.7), in an attempt topvoid further taxonomic
confusion, and because the type specimen of H.
longispina is lost, we propose that the name H.
longispina be suppressed and H. liebusi adopted as the
valid name for these middle Eocene morphotypes.
Shokhina illustrated a large number of H. liebusi
syntypes that clearly represent the range of morphology
exhibited by this taxon. Since no holotype was designated we herein select Shokhina's (1937) text-fig.
2 as the lectotype of H. liebusi and text fig. 8 as the
paralectotype (PI. 8.9, Figs. 1-2). Blow (1 979) did not
recognise H. liebusi and instead assigned Shokhina's
(1937) morphotypes either to H. mexicana sensu lato
or H. dumblei. We consider H. liebusi to be a useful
species that is morphologically and stratigraphically
intermediate between H. mexicana and H. dumblei.
PHYLOGENETIC RELATIONSHIPS.-
Hantkenina
liebusi evolved from H. mexicana by forward
progression of the tubulospine and closer appression of
the chambers. It gave rise to H. lehneri and H. dumblei by radial increase in chamber length and relaxation of
the coiling to increase the number of chambers. In H.
lehneri the chambers became elongate and the stellate
peripheral outline was retained, whereas in H. dumblei
the tubulospines moved forwards to span the anterior
suture, and the periphery was smoothed.
STRATIGRAPHIC RANGE.-
Middle Eocene Zones mid E8 to basal E13.
GEOGRAPHIC DISTRIBUTION.-
Global
distribution in the mid to low latitudes. Absent from
high southern or northern latitudes (Stott and Kennett,
1990; Huber, 1991).
STABLE ISOTOPE PALEOBIOL0GY.-
Hantkenina
liebusi has more positive ň18O and negative ň13C than
Morozovelloides and has a similar isotopic signature to
Turborotalia frontosa suggesting a relatively cold, deep
habitat (Pearson et al, 1993, 2001; Coxall and others,
2000). |
| Systematics: |
35
Ordo Foraminiferida
Superfamilia Globigerinaceae
Familia Guembelitriidae
Genus Jenkinsina
Species Hantkenina liebusi
Species Jenkinsina columbiana
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| Synonym list: |
Pearson et al. (2006):
1875 Siderolina kochi Hantken. - Hantken : p.79 pl. 16; fig. 1 [upper
Eocene, Clavulina-szaboi beds, Porva, Hungary]
1911 Pullenia kochi Liebus. - Liebus : p.942 pl. 11; fig. 9-10 [middle
Eocene, Yugoslavia]
1924 Hantkenina kochi Hantken. - Cushman : p.2 pl. 2; fig. 1a-c (reillustration
from Hantken, 1875)
? 1924 Hantkenina longispina Cushman. - Cushman : p.2 pl. 2; fig. 4 [Eocene, Rio Tuxpan, Tierra Colorado, Mexico]
1937 Hantkenina liebusi Shokhina. - Shokhina : p.427 pl. 2; fig. 2-3 [middle Eocene, Ilskaya, northern Caucasus, Russia]
? 1939 Hantkenina longispina Cushman. - Rey : p.323 pl. 22; fig. 1-2 [middle Eocene, Lutétien supérior,
Nummulitique de Rharb, Morocco]
1950 Hantkenina (Applinella) liebusi Shokhina. - Brönnimann : p.410 pl. 56; fig. 1, 2, 18, 19, 22, 23 [upper Eocene, San Fernando
Cp., Hospital Hill Fm., Trinidad]
1950 Hantkenina (Aragonella) aff. mexicana Cushman. - Brönnimann : p.407 pl. 55; fig. 9-10, 14 [lower and
middle Eocene, fig. 9, Oceanic Fm., Barbados; fig. 10,
Navet Fm., San Fernando, Trinidad; fig. 14, Navet Fm.,
Trinidad]; [Not Cushman, 1924]
1953 Hantkenina liebusi Shokhina. - Subbotina : p.148 pl. 1; fig. 11 [middle Eocene,
northern Caucasus, Russia]
1968 Hantkenina cf. liebusi Shokhina. - Raju : p.290 pl. 1; fig. 6 [Globigerina frontosa zone, Karaikal, Cauvery Basin,
southern India]
? 1975 Hantkenina longispina Cushman. - Stainforth et al. : p.203 pl. 64; fig. 1-6 [figs. 4-6, northeastern
Gulf of Mexico]
1993 Hantkenina liebusi Shokhina. - Pearson et al. : p.128 pl. 3; fig. 10 [middle Eocene, Zones P11-P12,
ODP Site 523, south Atlantic Ocean]
2006 Hantkenina liebusi Shokhina. - Pearson et al. : p.243 pl. 8.9; fig. 1-20 (Pl. 8.9, Fig. 1: illustration of lectotype [selected
herein] reillustrated from Shokhina, 1937)
(PI. 8.9, Fig. 2: illustration of paralectotype [selected
herein] reillustrated from Shokhina, 1937)
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| Specimen: |
Geological Research and Prospecting Institute - Leningrad, Inventory number: 60X No. 372
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| References: |
Hantken,V.M.. (1875):
Die fauna der Clavulina szahoi-schichten. 1. Teil: Foraminifera . Mitteilungen aus dem Jahrbuch der ungarischen Geologischen Anstalt, Budapest Vol. 4 p. 1-93
Liebus,A.. (1911):
Die foraminiferen fauna der mitteleocaenen Mergle von Norddal matlen . Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften in Wien, Mathematisch-Naturwissenschaftliche Klasse Vol. 120 p. 805-956
Cushman,J.A. (1924):
A new genus of Eocene foraminifera . Proceedings of the U.S. National Museum Vol. 66 p. 1-4
Shokhina,V.A.. (1937):
The genus Hantkenina and its stratigraphical distribution in the north Caucasus: Problems of Palaeontology. , Publications of the Laboratory of Palaeontology2-3 p. 425-441
Rey,M. (1939):
Distribution stratigraphique des Hantkenina dans le Nummulitique du Rharb (Maroc) . Bulletin de la Société Géologique de France Vol. 5 p. 321-341
Thalmann,H.E. (1942):
Foraminiferal genus Hantkenina and its subgenera . American Journal of Sciences Vol. 240 p. 809-823
Brönnimann,P. (1950):
The Genus Hantkenina Cushman in Trinidad and Barbados, B.W.I. . Journal of Paleontology Vol. 24 p. 397-420
Subbotina,N.N. (1953):
Fossil foraminifera of the USSR. Globigerinidae, Hantkeninidae and Globorotaliidae [in Russian] . Trudy Vsesoyuznogo Neftyanogo Nauchno-Isledovatelskogo Geologo-Razvedochnogo Instituta (VNIGRI), Novaya Seriya Vol. 76 p. 1-296
Ramsay,W.R. (1962):
Hantkenininae in the Tertiary rocks of Tanganyika . Contributions from the Cushman Foundation for Foraminiferal Research Vol. 13 p. 79-89
Raju,D.S.N. (1968):
Eocene-Oligocene planktonic foraminiferal biostratigraphy of Cauvery basin, South India . Memoires of the Geological Society of India Vol. 2 p. 286-289
Stainforth,R.M.; Lamb,J.L.; Luterbacher,H.P.; Beard,J.H. and Jeffords,R.M. (1975):
Cenozoic planktonic foraminiferal zonation and characteristics of index forms . Paleontological ContributionsArticle 62 p. 425
Pearson,P.N.; Shackleton,N.J. and Hall,M.A. (1993):
The Stable Isotope Paleoecology of Middle Eocene Planktonic Foraminifera and Multi-species integrated Isotope Stratigraphy . Journal of Foraminiferal Research Vol. 23 p. 123-140
Pearson,P.N.; Olsson,R.K.; Hemleben,C.; Huber,B.T. and Berggren,W.A. (2006):
Atlas of Eocene Planktonic Foraminifera. p. 1-513
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