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Morozovelloides lehneri Cushman & Jaris 1929 from: Pearson, P.N.Olsson, R.K.Hemleben, C.Huber, B.T.Berggren, W.A. (2006): Atlas of Eocene Planktonic Foraminifera. p. 1-513
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Species Morozovelloides lehneri Cushman & Jaris 1929



Diagnosis / Definition:
Pearson et al. (2006):
DESCRIPTION. Type of wall: Muricate, normal perforate, with concentrations of muricae around periphery and umbilicus and largely smooth chamber surfaces in between. Test morphology: Low trochospiral, elongateoval, strongly lobulate, normal perforate; early chambers muricate, later chambers essentially smooth, with scattered muricae around umbilical region; chambers weakly inflated, compressed; distinct blade-like or fimbriate peripheral muricocarina; 5-9 radially elongate, slightly inflated subtriangular to wedge-shaped chambers on umbilical side, flattening towards peripheral margin; intercameral sutures weakly curved on early portion of final whorl, essentially straight, radial and depressed on later part; umbilicus closed to narrowly open (in older forms), uniformly open and shallow in younger forms, rimmed by elevated circumumbilical chamber tips; aperture a low, umbilical-extraumbilical slit extending to periphery; approximately 12- l4 chambers arranged in 2 1/2 to 3 whorls (early chambers often obscured by muricae); sutures depressed, straight; periphery rimmed with a thick, blade-like to fimbriate muricocarina which, in the younger/later part of the test exhibits a tendency to become discontinuous; in edge view low biconvex trochospire, ante penultimate chambers gently sloping, raised circumumbilical tips of earlier chambers of last whorl projecting outwards. Size: Holotype: 0.36 mm; thickness/width: 0.15 mm (Cushman and Jarvis, 1929, p. 17).
Discussion / Comments:
Pearson et al. (2006):
DISTINGUISHING FEATURES.- Characterized by its low-biconvex, very thin, and essentially smooth, glossy test, its elongate, wedge-shaped chambers, straight to weakly curved, radial sutures on both the umbilical and spiral sides, and distinct blade-like to fimbriate peripheral muricocarina. It grades into Morozovelloides coronatus in the lower part of its range, where it is distinguished by the more elongate and more numberous chambers and tendency for crowding of the last one or two chambers. DISCUSSION.- The holotype of Morozovelloides lehneri is illustrated by SEM for the first time on P1. 10.4, Figs. 1-3. It is not a very well-preserved specimen but it shows the key morphological features of the taxon. With its smooth, translucent (when well preserved) elongate-petaloid test, radially elongate chambers and blade-like to fimbriate muricocarina, this is one of the most distinctive of the Eocene species. We use it to typify the new genus Morozovelloides. Blow (1979, p. 1002) pointed to the phylogenetic trend observed in this morphotype of a gradual enlargement of the umbilicus and ontogenetic disruption/discontinuity in the peripheral muricocarina in younger/later chambers of the final whorl, which may be of stratigraphic utility- In our investigation of well- preserved populations from Tanzania, we have observed a recurrent morphotype with chevron-pointed chamber ends (see Plate 10.4, Fig. 15) but it is unclear whether this variety is of stratigraphic utility. PHYLOGENETIC RELATIONSHIPS-. This morphotype evolved from Morozovelloides coronatus the early middle Eocene by radial elongation of the chambers, further compression of the test and development of a thickened, blade-like and/or fimbriate muricocarina. STRATIGRAPHIC RANGE.- Uppermost Zone E8- E12. GEOGRAPHIC DISTRIBUTION.- Cosmopolitan in the tropics and subtropics. STABLE ISOTOPE PALEOBIOL0GY.- Morozo- velloides lehneri has consistent oxygen and carbon isotopic ratios indicative of a shallow mixed layer habitat (Boersma and others, 1987; Pearson and others, 2001).
Systematics:

35
 Ordo Foraminiferida
  Superfamilia Globigerinaceae
   Familia Truncorotaloididae
    Genus Morozovelloides
     Species Morozovelloides lehneri
Synonym list:
Pearson et al. (2006):
1929 Globorotalia lehneri Cushman & Jarvis. - Cushman & Jarvis : p.17 pl. 3; fig. 16a-c [middle Eocene, Lower marl, Trinidad]
1949 Globorotalia lehneri Cushman & Jarvis. - Cushman & Bermudez : p.32 pl. 6; fig. 7-9 [San Luis Fm., Oriente Province, C u b a ]
p 1957 Globorotalia lehneri Cushman & Jarvis. - Bolli : p.169 pl. 38; fig. 11a, b [Porticulasphaera mexicana Zone, Navet Fm., Trindad]
1961 Pseudogloborotalia lehneri Cushman & Jarvis. - Bermudez : p.1345 pl. 16; fig. 9 (2 fig.) [middle Eocene, Oriente Province, Cuba]
1962 Globorotalia lehneri Cushman & Jarvis. - Hofker : p.119 fig. 52a-c [middle Eocene Zone P 12, Guayabal Fm., Mexico]
1963 Globorotalia lehneri Cushman & Jarvis. - Aubert : p. 62, 63 pl. 5; fig. 1a-c [upper Lutetian, Jebel Si- Ameur-el Hadi, Morocco]
1969 Globorotalia lehneri Cushman & Jarvis. - Blow : p.363 pl. 50; fig. 1 [Zone P13, Zanzibar Well No. l, Island of Zanzibar, Tanzania]
1971 Globorotalia lehneri Cushman & Jarvis. - Postuma : p.198 fig. on p. 199 [middle Eocene, Dyr el Kef, Tunisia]
1972 Globorotalia lehneri Cushman & Jarvis. - Samuel : p.193 pl. 49; fig. 2a-c [middle Eocene Bakony Mountains, Borehole MO-l, Hungary]
1975 Globorotalia lehneri Cushman & Jarvis. - Toumarkine : p.738 pl. 2; fig. 13, 14 [G. lehneri Zone, DSDP Site 3 13, South Atlantic]
1979 Globorotalia (Morozovella) lehneri Cushman & Jarvis. - Blow : p. 1002, 1003 pl. 50, fig. 1 (from Part l, 1969; see above); pl. 188, fig. 1-10 [Zone P12, type locality of Guayabal Fm., Arroyo Gallo, Mexico]; pl. 251, fig. 3,4 [Zone P12, Mafia well No. 1, Mafia Island, East Africa; incorrectly listed as plate 197 by Blow, 1979:1003]
1985 Morozovella lehneri Cushman & Jarvis. - Toumarkine & Luterbacher : p.131 pl. 3 1. 1 (holotype refigured), 3 1.2(Cushman and Bermudez, 1949, refigured), 3 1.3 (Bolli, 1957, pl. 38, figs. l l a,b refigured), 3 1.46 [M. lehneri Zone, El Datil Fm., Punta Mosquito, Margarita Island, Venezuela], 3 1. 7-8 [M. lehneri Zone, Baluchistan, Pakistan], 3 1.9-13 [M. lehneri Zone, Guayabal Fm., Tampico Embayment, Mexico]
2006 Morozovelloides lehneri Cushman & Jaris. - Pearson et al. : p.339 pl. 10.4; fig. 1-16 (Pl. 10.4, Figs. 1-3: new SEMs of holotype of Globorotalia lehneri Cushman and Jarvis)
Specimen:
Cushman Collection - Smithsonian Museum of Natural History, Washington, D.C., Inventory number: CC 10068
References:

Cushman,J.A. and Jarvis,P.W. (1929):
New foraminifera from Trinidad . Contributions from the Cushman Foundation for Foraminiferal Research Vol. 5 p. 6-17

Cushman,J.A. and Bermudez,P.J. (1949):
Some Cuban species of Globorotalia . Contributions from the Cushman Foundation for Foraminiferal Research Vol. 25 p. 26-44

Bolli,H.M. (1957):
Planktonic Foraminifera from the Eocene Navet and San Fernando formations of Trinidad, B.W.I. . Bull. U.S. natl. Mus. Vol. 215 p. 155-172

Bermudez,P.J. (1961):
Contribucion al estudio de las Globigerinidea de la region Caribe-Antillana (Paleocene-Reciente).
In: Boletino Geologia (Venezuela), Special Publicacion Vol. 3

Hofker,J. (1962):
Änderung des Generationswechsels der Foraminiferen-Arten während des Periods ihres Dasiens . Neues Jahrbuch für Geologie und Palaontologie, Monatshefte Vol. 8 p. 316-329

Aubert,J.. (1963):
Les Globorotalia de la region prerifaine (Maroc septentrional) . Notes et Memoires du Service Geologique Maroc Vol. 22 p. 1-156

Blow,W.H. (1969):
Late middle Eocene to Recent planktonic foraminiferal biostratigraphy.
In: Proceedings of the First International Conference on Planktonic Microfossils, Geneva 1967 Vol. 1 Eds: Bronnimann, P.Renz, H.H. p. 199-422

Postuma,J.A. (1971):
Manual of Planktonic Foraminifera. p. 422 pp

Samuel,O.. (1972):
New species of planktonic foraminifers from the Paleogene of the west Carpathian in Slovakia (Czechoslovakia) . Zbornik Geologick"ck Vied Západné Karpaty Vol. 17 p. 165-221

Toumarkine,M. (1975):
Middle and Late Eocene planktonic foraminifera from the northwestern Pacific, Leg 32 of the Deep Sea Drilling Project.
In: Initial Reports of the Deep Sea Drilling Project Vol. 32 Eds: Larson, R.L.Moberly, R. p. 735-751

Blow,W.H. (1979):
The Cainozoic Globigerinida. 3 Vols p. 1413 pp

Toumarkine,M. and Luterbacher,H.P. (1985):
Paleocene and Eocene Planktic Foraminifera.
In: Plankton Stratigraphy p. 87-154

Pearson,P.N.; Olsson,R.K.; Hemleben,C.; Huber,B.T. and Berggren,W.A. (2006):
Atlas of Eocene Planktonic Foraminifera. p. 1-513

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