Chiloguembelina ototara Finlay 1940 from: Pearson, P.N.Olsson, R.K.Hemleben, C.Huber, B.T.Berggren, W.A. (2006): Atlas of Eocene Planktonic Foraminifera. p. 1-513 . |
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Species Chiloguembelina ototara Finlay 1940 |
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Diagnosis / Definition: |
Pearson et al. (2006):
DESCRIPTION.
Type of wall: Microperforate, uniformly finely
pustulose, lacking pore mounds or costae.
Test morphology: Test short to somewhat
elongate, moderately to rapidly expanding, subtriangular
in outline, periphery rounded rather than compressed;
chambers biserial, increasing moderately to rapidly in
size, usually 11-12, and sometimes up to 15 in adult
specimens; sutures depressed, perpendicular to slightly
oblique to growth axis; aperture a moderately narrow to
broad symmetrical arch centered or slightly off-center of the base of the final chamber, extending half-way up
the final chamber face, bordered on one side by a narrow
lip that thickens away from its attachment point on the
chamber face.
Size: Paratype USNM 689 102: length, 0.17 mm,
width 0.10 mm, breadth 0.07 mm; hypotypes up to 0.23
mm length, 0.9 mm breadth. |
Discussion / Comments: |
Pearson et al. (2006):
DISTINGUISHING FEATURES.-
Distinguished from
C. cubensis by the absence of striae on the wall surface
and the shorter, more strongly tapering test, and from S.
martini by its more inflated chambers and lower apertural
arch.
DISCUSSION.-
Hornibrook (1990) was the first to
draw attention to the significant difference in wall texture
between topotypes of C. cubensis, which are striate, and
topotypes of C. ototara, which have pustules that are
randomly distributed on the test surface rather than striae.
These differences are apparent in SEM images but they
are difficult to discern under the light microscope.
Guembelina cubensis Palmer var. heterostoma
Bermudez and Guembelina venezuelana Nuttall var.
rugosa Parr may be prior synonyms but their wall
textures are currently undetermined.
Guembelina multicellaris Hussey is considered
synonymous with C. ototara based on SEM observation
of the multicellaris holotype (PI. 16.3, Figs. 9,10), which
reveals an elongate test with inflated chambers, a low,
interiomarginal aperture, and pustules rather than striae
on the wall surface.
PHYLOGENETIC RELATIONSHIPS.-
Probably
descended from Chiloguembelina crinita during the early
middle Eocene.
STRATIGRAPHIC RANGE.-
Middle Eocene Zone E9
through lower Oligocene Zone 01. The lowest and
highest occurrences of this species are difficult to
determine because of its morphologic similarity to its
presumed ancestor, C. crinita, and its likely descendant,
C. cubensis. A lowermost Oligocene (Zone 01)
extinction of this species was recorded in New Zealand
(Hornibrook, 1990) and ODP Sites 806 and 807 on
Ontong Java Plateau (Resig, 1993).
GEOGRAPHIC DISTRIBUTION.-
Cosmopolitan.
STABLE ISOTOPE PALEOBIOLOGY.-
Stable
isotope data obtained for upper Eocene C. ototara
(Barrera and Huber, 1991; identified as Chiloguembelina
spp.) suggest that it lived near the surface mixed layer
at high southern latitudes.
REPOSITORY.-
Holotype and paratypes deposited in
the Geological and Nuclear Research Institute, Lower
Hutt, New Zealand; paratype (USNM 689102) also
deposited at the Smithsonian Museum of Natural History,
Washington, D.C. |
Systematics: |
35 Ordo Foraminiferida
Superfamilia Globigerinaceae
Familia Chiloguembelinidae
Genus Streptochilus
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Synonym list: |
Pearson et al. (2006):
1940 Guembelina ototara Finlay. - Finlay : p.453 pl. 63; fig. 50-52 [Oamaru Diatomite, latest Eocene (Runangan Stage) South
Island, New Zealand]
1985 Chiloguembelina cubensis Palmer. - Jenkins & Srinivasan : p.807 pl. 1; fig. 2 [upper Eocene Globigerina linaperta Zone, DSDP
Site 592, Lord Howe Rise]
1990 Chiloguembelina ototara Finlay. - Hornibrook : pl. 1; fig, 12-16 [Oamaru Diatomite, latest Eocene (Runangan
Stage), South Island, New Zealand]
1991 Chiloguembelina cubensis Palmer. - Huber : p.440 pl. 6; fig. 8 [Zone AP13, lower Oligocene, ODP Hole 738B,
Kerguelen Plateau, southern Indian Ocean];[
Not Palmer,
1934]
1993 Chiloguembelina ototara Finlay. - Resig : pl. 1; fig. 9 [upper Eocene, ODP Site 807, Ontong Java Plateau,
western Pacific Ocean]
2006 Chiloguembelina ototara Finlay. - Pearson et al. : p.480 pl. 16.3; fig. 9, 10, 13-22 (Pl. 16.3, Figs. 9, 10: new SEMs of holotype of
Guembelina multicellaris Hussey)
(Pl. 16.3, Figs. 13, 14: new SEMs of paratype of
Guembelina ototara Finlay)
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Specimen: |
Smithsonian Museum of Natural History, Washington, D.C., Inventory number: USNM 689102
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References: |
Bermudez,P.J. (1937): Nuevas especies de foraminiferos del Eoceno, de Cuba . Mem. Soc. Cubana Hist. Nat. Vol. 11 p. 137-150
Parr,W.J. (1938): Upper Eocene foraminifera from the deep borings in King's Park, Perth, Western Australia . J. R. Soc. Western Australia Vol. 24 p. 69-101
Finlay,H.J. (1940): New Zealand Foraminifera; key species in stratigraphy . Transactions of the Royal Society of New Zealand Vol. 69
Jenkins,D.G. (1971): New Zealand Cenozoic Planktonic Foraminifera . New Zealand Geological Survey Paleontological Bulletin Vol. 42
Jenkins,D.G. and Srinivasan,M.S. (1985): Cenozoic planktonic foraminifera from the Equator to the Sub-Antarctic of the Southwest Pacific. In: Initial Reports of the Deep Sea Drilling Project Vol. 90 Eds: Kennett, J.P.von der Borchand others p. 795-834
Hornibrook,N. (1990): Chiloguembelina cubensis (Palmer) and C. ototara (Finlay), in New Zealand . Journal of Foraminiferal Research Vol. 20 p. 368-371
Huber,B.T. (1991): Paleogene and early Neogene planktonic foraminifer biostratigraphy of Sites 738 and 744, Kerguelen Plateau (southern Indian Ocean). In: Proceedings of the Ocean Drilling Program, Scientific Results Vol. 119 Eds: Leckie, R.M.Sigurdsson, H.Acton, G.D.Draper, G. p. 427-449
Resig,J.M.. (1993): Cenozoic stratigraphy and paleoceanography of biserial planktonic foraminifers, Ontong Java Plateau . In: Proceedings of the Ocean Drilling Program, Scientific Results Vol. 130 Eds: Berger, W.H.Kroenke, L.W..Mayer, L.A..and others p. 231-244
Pearson,P.N.; Olsson,R.K.; Hemleben,C.; Huber,B.T. and Berggren,W.A. (2006): Atlas of Eocene Planktonic Foraminifera. p. 1-513
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