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Morozovella velascoensis Cushman 1925 from: Pearson, P.N.Olsson, R.K.Hemleben, C.Huber, B.T.Berggren, W.A. (2006): Atlas of Eocene Planktonic Foraminifera. p. 1-513
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Species Morozovella velascoensis Cushman 1925



Discussion / Comments:
Toumarkine & Luterbacher (1985):
Typical representatives of M. velascoensis are close to their predecessor M. conicotruncata to which they are linked by a series of transitional forms. Characteristic for the species are the raised and beaded intercameral sutures on the spiral side and the heavy spines on the periphery which fuse to form a heavy 'keel' (muricocarina). Such spines are also present on the umbilical chmaber tips.
Van Eijden & Smit (1991):

Pearson et al. (2006):
DISCUSSION.- This taxon was discussed in the Atlas of Paleocene Planktonic Foraminifera (Olsson and others, 1999). STRATIGRAPHIC RANGE.- P3b to Zone E2.
Systematics:

1
 Superregnum Eukaryota
  Regnum Protoctista
   Phylum Ciliophora
    Subphylum Postciliodesmatophora
     Ordo Globigerinida
      Superfamilia Globorotaliaceae
       Superfamilia Nonionacea
        Familia Truncorotaloididae
         Genus Morozovella
          Species Morozovella velascoensis

15
  Classis Foraminifera
   Genus Morozovella
    Species Morozovella velascoensis

17
 Classis Foraminifera
  Genus Morozovella
   Species Morozovella velascoensis

19
  Ordo Foraminiferida
   Superfamilia Globorotaliaceae
    Familia Truncorotaloididae
     Genus Morozovella
      Species Morozovella velascoensis

32
 Ordo Foraminiferida
  Familia Globorotaliidae
   Genus Morozovella
    Species Morozovella velascoensis

35
  Ordo Foraminiferida
   Superfamilia Globigerinaceae
    Familia Truncorotaloididae
     Genus Morozovella
      Species Morozovella velascoensis
Synonym list:
Berggren & Norris (1997):
1925 Pulvinulina velascoensis Cushman. - Cushman : p. 19 pl. 3, figs. 5a-c (Velasco Shale Formation, San Luis Potosi, Tampico Embayment, E Mexico).
1926 Pulvinulina velascoensis Cushman. - Cushman : p. 608 pl. 21, figs. 9a,b (reillustration of holotype).
1927 Pulvinulina velascoensis Cushman. - Cushman : p. 169 pl. 27, figs. 7-9 (Velasco Shale, Mexico).
1928 Globorotalia velascoensis Cushman. - White : p. 281 pl. 398, figs. 2a-c (Verlasco Shale Formation, E Mexico).
1934 Globorotalia velascoensis Cushman. - Dampel : p. 28 pl. 4, figs. 2a-c (Emba region, SW Russia).
1945 Globorotalia velascoensis Cushman. - Applin & Jordan : p. 146 pl. 19, figs. 8a,b (subsurface, NW Florida).
1946 Globorotalia velascoensis Cushman. - Cushman & Renz : p. 47 pl. 8, figs. 13, 14 (lower Lizard Springs Formation, Trinidad).
1947 Globorotalia velascoensis Cushman. - Subbotina : p. 123 pl. 7, figs. 9-11 (G. velascoensis Zone, Sunzha River section, N Caucasus); pl. 9, figs. 21-23 (G. velascoensis Zone, Kuban River section, N Caucasus).
p 1953 Globorotalia velascoensis Cushman. - Hamilton : p. 231 pl. 30, figs.16-18 (Hess Guoyot, central equatorial Pacific); non pl. 30, fig. 23; non pl. 31, figs. 28, 29, 31; ? figs. 24, 30; LE ROY 1955, p. 33, pl. 3, figs. 1-3 (Maqfi secton, Egypt).
1953 Globorotalia velascoensis Cushman. - Subbotina : p. 216 pl. 19, figs. 1a-2c (Kuban River section, Foraminiferal Beds, Green Group, (Zone of conical globorotaliids= Morozovella caucasica (GLAESSNER); pl. 19, figs. 3a-4c (Gubs River section, Foraminiferal Beds, Kutais Horizon (F2), Barakaen region, N Caucasus, Zone of conical globorotaliids= Morozovella aragonensis (NUTTALL).
1956 Globorotalia velascoensis Cushman. - Haque : p. 181 pl. 24, figs. 2a-c (upper Paleocene, Nammal Gorge, Salt Range, Pakistan).
1957 Globorotalia velascoensis Cushman. - Bolli : p. 76 pl. 20, figs. 1-4 (G. pseudomenardii Zone, lower Lizard Sprngs Formation, Trinidad).
1957 Globorotalia velascoensis Cushman. - Loeblich & Tappan : p. 196 pl. 64, figs. la-2c (Velasco Formation, Tamaulipas, Mexico).
1959 Globorotalia velascoensis Cushman. - Nakkady : p. 462 pl. 4, figs. 4a-c (Um Elghanayem section, Western Desert, Egypt).
1959 Globorotalia velascoensis Cushman. - Hamilton & Rex : p. 794 pl. 252, figs. 18-20 (Sylvania
1960 Globorotalia velascoensis Cushman. - Bolli & Cita : p. 391 pl. 35, figs. 7a-c (G. pseudomenardii Zone, Paderno d'Adda section, N Italy).
1960 Pseudogloborotalia velascoensis Cushman. - Bermudez : p. 1349 pl. 16, figs. 1 la-b (4 figs.) (Velasco Station, Tampico-San Luis Potosi Railway, Mexico) (specimen USNM 639063 mislabeled as holotype of Pseudogloborotalia pasionensis by Bermudez).
1961 Globorotalia velascoensis Cushman. - Said & Kerdany : p. 330 pl. 1, figs. l0a-c (lower Esna Shale Formation, Western Desert, Egypt).
1962 Globorotalia (Truncorotalia) velascoensis velascoensis Cushman. - von Hillebrandt : p. 169 pl. 13, figs. 16-2l (Zone D=G. pusilla pusilla Zone (upper part), Reichenhall-Salzburg Basin, Bavaria, S Germany).
1963 Truncorotalia velascoensis velascoensis Cushman. - Gohrbandt : p. 59 pl. 5, figs. 7-9 (Salzburg basin, Austria).
1964 Globorotalia velascoensis Cushman. - Luterbacher : p. 681 text-figs. 92a-96c, 98a,b (G. velascoensis Zone, Ebano, Mexico); text-figs. 99a-c (G. velascoensis Zone), l00a-c (G. aequa Zone, Gubbio section, N Apennines, N Italy).
1964 Globorotalia velascoensis Cushman. - Said & Sabry : p. 386 pl. 2, figs. 9a-c (Esna Shale Formation, Gebel Aweina, Egypt).
1965 Globorotalia velascoensis Cushman. - Proto Decima & Zorzi : p. 32,33 pl. 4, figs, la-c ; pl. 5, figs. 3, 4 (G. pseudomenardii Zone, Molinetto di Pederobba section, W Trevignano, Italy; non pl. 4, figs. 2-7).
1966 Globorotalia velascoensis Cushman. - El Naggar : p. 246 pl. 20, figs. 3a-d; pl. 20, fig. 3 (G. velascoensis Zone, Oweina Shale Formation, Egypt).
1968 Truncorotaloides (Morozovella) velascoensis Cushman. - McGowran : pl. 2, fig. 1 (Velasco Shale Formation, Ebano, Mexico).
1970 Globorotalia velascoensis Cushman. - Samanta : p. 636 pl. 97, figs. 7-10 (G. pseudomenardii Zone, upper marlstone, Pondicherry Formation, Madras, S India).
1970 Globorotalia velascoensis Cushman. - Shutskaya : p.118-120 pl. 23, figs. 3a-c, pl. 24, figs. 5a-c (Ac. tadjikistanensis djanensis Subzone, Kachan Stage, Tarkhankut Peninsula, Crimea, Olenev Platform, Boring 210,517-527m); pl. 25, figs. 5a-c (same zone as above, but from boring 229, 637-641m); pl. 27, figs. 12a-c (Ac. acarinata Zone, Kachan Stage, Tarkhankut Peninsula, Crimea; pl. 29, figs. 8a-c (G. aequa Zone, Bakhchissarayan Stage, Tarkhankut Peninsula, Crimea).
1971 Globorotalia velascoensis Cushman. - Postuma : p. 218 7 text-figs, on p, 219 (Topotypes from Velasco Shale Formation, Tamalte Arroyo, Hacienda El Limon, San Luis Potosi, Mexico).
1971 Globorotalia (Morozovella) velascoensis velascoensis Cushman. - Jenkins : p. 107 pl. 9, figs. 214-216 (G. triloculinoides Zone, Waipawan Stage, Middle Waipara River section, New Zealand).
1975 Globorotalia velascoensis Cushman. - Luterbacher : p. 726 pl. 1, figs. 8a,b (G. pseudomenardii Zone, DSDP 32/305/14/1: 135- 138cm, Shatsky Rise, NW Pacific).
1977 Morozovella velascoensis Cushman. - Berggren : p. 232 Plate 6 (refigured specmens from the literature).
1979 Globorotalia (Morozovella) velascoensis velascoensis Cushman. - Blow : p. 1029 pl. 92, fig. 7 (Zone P4, DSDP6/47.2/9/3:70- 72cm); pl. 94, figs. 6-9; pl. 95, figs. 1,2; pl. 216, figs. 1-8; pl. 217,figs. 1-6 (Zone P5, Samples FCRM. 1670, Lindi area, Tanzania, E Africa); pl. 99, figs. 3,4 (Zone P5, DSDP 6/47.2/9/1: 64-66cm).
1983 Globorotalia velascoensis Cushman. - Pujol : p. 644 pl. 3, fig. 9 (Zone P5, DSDP 72/516F/84/5: 13-15cm; Rio Grande Rise, SW Atlantic).
1984 Globorotalia (Morozovella) velascoensis velascoensis Cushman. - Belford : p. 10 pl. 1, fig. 2; pl. 19, figs. 1-12 (upper Paleocene, Wabag Sheet area, Papua, New Guinea).
1985 Morozovella velascoensis Cushman. - Toumarkine & Luterbacher : p. 110 text-figs. 13 (11: [reillustration of BOLLI 1957a, pl. 20, figs. 1-3]; 12 (reillustration of holotype from Cushman 1925, pl. 3, figs. 5a-c]).
1988 Morozovella velascoensis Cushman. - Pandey & Ravindram : p. 173, 174 pl. 13, figs. 9 (sample 875-880m); 10 (sample at 890- 895m); 11-13 (core at 985,6-995m) (Zone P4, KRI-A well, Krishna- Godavary Basin, offshore E India).-BRALOWER et al 1995, p. 844, Figure 2 (1,2) (Zone P4/5, ODP143/865C/12H/4-10-12cms; Allison Guyot, Mid-Pacific Mountains, W Equatorial Pacific).
1997 Morozovella velascoensis Cushman. - Berggren & Norris : p. 101 Plate 15 Figures 10-15,16-18,22
Toumarkine & Luterbacher (1985):
1925 Pulvinulina velascoensis Cushman. - Cushman : p.19 pl 3 fig 5 (type reference)
1985 Morozovella velascoensis Cushman. - Toumarkine & Luterbacher : p.109 figs. 13.11-12
Olsson et al. (1999):
1925 Pulvinulina velascoensis Cushman. - Cushman : p. 19 pl. 3, figs. 5a-c (Velasco Fm., San Luis Potosi, Tampico Embayment, eastern Mexico).
1928 Globorotalia velascoensis Cushman. - White : p. 281 pl. 398, figs. 2a-c (Velasco Shale Fm., E Mexico)
1946 Globorotalia velascoensis Cushman. - Cushman & Renz : p. 47 pl. 8, figs. 13, 14 (lower Lizard Springs Fm., Trinidad)
1947 Globorotalia velascoensis Cushman. - Subbotina : p. 123 pl. 7, figs. 9-11 (G. velascoensis Zone, Sunzha River section, N Caucasus); pl. 9, figs. 21-23 (G. velascoensis Zone, Kuban River section, N Caucasus)
1953 Globorotalia velascoensis Cushman. - Le Roy : p. 33 pl. 3, figs. 1-3 (Maqfi secton, Egypt)
1956 Globorotalia velascoensis Cushman. - Haque : p. 181 pl. 24, figs. 2a-c (upper Paleocene, Nammal Gorge, Salt Range, Pakistan)
1957 Globorotalia velascoensis Cushman. - Bolli : p. 76 pl. 20, figs. 1-4 (G. pseudomenardii Zone, lower Lizard Springs Fm., Trinidad)
1957 Globorotalia velascoensis Cushman. - Loeblich & Tappan : p. 196 pl. 64, figs. 1a-2c (Zone P4, Velasco Fm., Tamaulipas, Mexico)
1960 Globorotalia velascoensis Cushman. - Bolli & Cita : p. 391 pl. 35, figs. 7a-c (G. pseudomenardii Zone, Paderno d'adda section, N Italy)
1961 Globorotalia velascoensis Cushman. - Said & Kerdany : p. 330 pl. 1, figs. 10a-c (lower Esna Shale Fm., Western Desert, Egypt)
1961 Pseudogloborotalia velascoensis Cushman. - Bermudez : p. 1349 pl. 16, figs. 11a-b (4 figs.) (Velasco Station, Tampico-San Luis Potosi Railway, Mexico)
1962 Globorotalia (Truncorotalia) velascoensis velascoensis Cushman. - von Hillebrandt : p. 169 pl. 13, figs. 16-21 (Zone D=G. pusilla pusilla Zone (upper part), Reichenhall-Salzburg Basin, Austro-German border).
1968 Truncorotaloides (Morozovella) velascoensis Cushman. - McGowran : pl. 2, fig. 1 (Velasco Fm., Ebano, Mexico)
1970 Globorotalia velascoensis Cushman. - Shutskaya : p. 118-120 pl. 23, figs. 3a-c, pl. 24, figs. 5a-c (Ac. tadjikistanensis djanensis Subzone, Kachan Stage, Tarkhankut Peninsula, Crimea, Olenev Platform, Boring 210, 517-527 m); pl. 25, figs. 5a-c (same zone as above, but from boring 229, 637-641 m); pl. 27, figs. 12a-c (Ac. acarinata Zone, Kachan Stage, Tarkhankut Peninsula, Crimea; pl. 29, figs. 8a-c (G. aequa Zone, Bakhchissarayan Stage, Tarkhankut Peninsula, Crimea)
1971 Globorotalia (Morozovella) velascoensis velascoensis Cushman. - Jenkins : p. 107 pl. 9, figs. 214-216 ( G. triloculinoides Zone, Waipawan Stage, Middle Waipara River section, New Zealand
1975 Globorotalia velascoensis Cushman. - Luterbacher : p. 726 pl. 1, figs. 8a,b (G. pseudomenardii Zone, DSDP Site 305/14/1: 135-138 cm, Shatsky Rise, NW Pacific Ocean)
1979 Globorotalia (Morozovella) velascoensis velascoensis Cushman. - Blow : p. 1029 pl. 92, fig. 7 (Zone P4, DSDP Hole 47.2/9/3:70-72 cm, Shatsky Rise, NW. Pacific Ocean); pl. 94, figs. 6-9; pl. 95, figs. 1,2; pl. 216, figs. 1-8; pl. 217, figs. 1-6 (Zone P5, Samples FCRM. 1670, Lindi area, Tanzania, E Africa); pl. 99, figs. 3,4 (Zone P5, DSDP Hole 47.2/9/1: 64-66 cm, Shatsky Rise, NW. Pacific Ocean)
1983 Globorotalia velascoensis Cushman. - Pujol : p. 644 pl. 3, fig. 9 (Zone P5, DSDP Hole 516F/84/5: 13-15 cm; Rio Grande Rise, SW Atlantic Ocean)
1984 Globorotalia (Morozovella) velascoensis velascoensis Cushman. - Belford : p. 10 pl. 1, fig. 2; pl. 19, figs. 1-12 (upper Paleocene, Wabag Sheet area, Papua, New Guinea)
1985 Morozovella velascoensis Cushman. - Toumarkine & Luterbacher : p. 110 text-figs. 13 (11: (reillustration of Bolli, 1957a, pl. 20, figs. 1-3); 12 (reillustration of holotype from Cushman, 1925, pl. 3, fgs. 5a-c)
Van Eijden & Smit (1991):
1925 Pulvinulina velascoensis Cushman. - Cushman : p.19 pl. 3, fig. 5
1991 Morozovella velascoensis Cushman. - Van Eijden & Smit : p.114
Pearson et al. (2006):
2006 Morozovella velascoensis Cushman. - Pearson et al. : p.348 pl. 11.2; fig. 1-3
Stratigraphy - absolute ages:
FAD: 60 ± 0 [Ma], Berggren et al. (1995) DSDP Leg 43 Site 384
LAD: 54.7 ± 0 [Ma], Berggren et al. (1995)
References:

Cushman,J.A. (1925):
Some new foraminfera from the Velasco Shale of Mexico . Contributions from the Cushman Foundation for Foraminiferal Research Vol. 1 p. 18-23

Cushman,J.A. (1926):
The Foraminifera of the Tampico embayment area . Bulletin American Association Petroleum Geologists Vol. 10(6) p. 581-612

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New and interseting foraminifera from Mexico and Texas . Contributions from the Cushman Foundation for Foraminiferal Research Vol. 3 p. 111-119

White,M.P. (1928):
Some index foraminifera of the Tampico embayment of Mexico, Part 2 . Journal of Paleontology Vol. 2(4) p. 280-313

White,M.P. (1928):
Some index foraminifera of the Tampico embayment of Mexico, Part 1 . Journal of Paleontology Vol. 2(3) p. 177-215

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Foraminifera from the upper Cretaceous of the Karaton field (Emba region). p. 33

Applin,E.R. and Jordan,L. (1945):
Diagnostic Foraminifera from subsurface formations in Florida . Journal of Paleontology Vol. 19(2) p. 129-148

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Reference not given in Berggren and Norris, 1997

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Atlas of Eocene Planktonic Foraminifera. p. 1-513

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