Pachydiscus (Pachydiscus) gollevillensis d'Orbigny 1850 from: Kennedy, W.J.. (1986): The ammonite fauna of the Calcaire à Baculites (Upper Maastrichtian) of the Cotentin Peninsula (Manche, France) . Palaeontology Vol. 29(1) p. 25-83
|Notice: This catalogue page may contain unedited data.
Species Pachydiscus (Pachydiscus) gollevillensis d'Orbigny 1850
|Diagnosis / Definition:
Cobban & Kennedy (1995):
USNM 463107, a phosphatic internal mold of a
phragmocone, has maximum preserved whorl height of 23 mm.
Coiling appears to have been moderately involute, with small,
shallow umbilicus; umbilical wall low and rounded. Whorl
breadth to height ratio 0.7, with greatest breadth just outside
umbilical shoulder. Inner flanks broadly rounded, mid- to outer
flanks flattened and convergent, venter very broadly rounded.
Four weak prorsiradiate bullae perched on umbilical shoulder
give rise to low, weak, prorsiradiate ribs that efface across middle
of flanks. In places, ribs appear to bifurcate. Intercalated ribs
arise on outer flank, where they are very weak; they strengthen
markedly over ventrolateral shoulder before weakening somewhat
at midventer and becoming interrupted by marked siphuncular
groove. Seventeen ventral ribs correspond to four
bullae. Suture complex and typical for genus.
Types. This is a Prodrome species, introduced by d'Orbigny in 1850 as nomen novum for Ammonites lewesiensis d'Orbigny (not Mantell): '*17. Gollevillensis, d'Orb. 1847. A. lewesiensis, d'Orb., 1842. Paleout. franc., Terr. cret., pI. 101 et 102, fig. 1 (non Sowerby). France, Golleville, Fresville (Manche)' (d'Orbigny 1850, p. 212). In Paleontologie Francalse d'Orbigny (1841, p. 336, pI. 101; pl. 102, figs. 1 and 2) notes that de Gerville had found the species at Golleville and Fresville.
The de Gerville Collection, formerly at Caen, was destroyed during the Second World War, while there are no specimens from either of the above localities listed in the catalogue of the d'Orbigny Collection under A. gollevillensis or lewesiensis. De Gerville sent ammonites from the Calcaire a Baculites to several of his contemporaries; there are, for instance, specimens labelled in his distinctive style in the Museum für Natürkunde, Berlin, Naturhistorisches Museum, Vienna, and the British Museum (Natural History). In the case of material sent to James Sowerby, one was described as A. constrictus in 1817, so dating the transaction some twenty-four years earlier at minimum than publication of Paleontologie Francalse and rendering it unlikely (but not impossible) that the associated Pachydiscus in Sowerby's Collection were studied by d'Orbigny and are thus surviving syntypes. Because d'Orbigny's protographs (reproduced here as text-fig. 2) represent at least two species, and because the original of his plate 101 is sufficiently idealized to be interpretable as either P. gollevillensis of authors or P. valognensis (Spath, 1922) for those who believe these species to be distinct, neotype designation is desirable. BMNH C38179 (PI. 1, figs. 1-3) is so designated in the interest of nornen¬clatural stability; the specimen being from Fresville and one of the specimens sent to James Sowerby by de Gerville.
Material. BMNH 50135 from Valognes, the holotype by monotypy of Parapachydiscus valognensis Spath, 1922a (PI. 2, figs. 1-3); BMNH C38178 and C70651 (internal and external moulds) from Fresville (ex J. Sowerby, ex de Gerville Collection); MNHP unreg. (de Vibraye Collection) from Fresville; SP 19 from Golleville (the original of Seunes 1891, pI. 14 (5), fig. 3; the repository given by Seunes is an error); EMP unreg., two specimens from Fresville or Valognes (the originals ofSeunes 1891, pI. 14 (5), figs. 1 and 2; the repository given by Seunes is an error); SP unreg. from Fresville (the original of de Grossouvre 1894, pI. 31, fig. 9); FSR 12, a juvenil. from Fresville (Seunes Collection); MNHH 5856, from Port Filiolet, Picauville; MHNG, two unreg. specimens (ex Pictet Collection) from Fresville; MNB no, Hi, from Valognes (ex de Gerville Collection).
Description. All specimens are internal moulds of phragmocones. No body-chambers known. Coiling moderately involute, with small, shallow umbilicus comprising around 25 % of diameter.
Umbilical walllow and rounded, whorl section compressed (whorl breadth to height ratio ranges from 0•67 to 0,75) with greatest breadth weil below mid-fiank (text-fig. 3p, R). Inner fianks broadly rounded, outer fianks fiattened and convergent, ventrolateral shoulders broadly rounded, venter flattened.
Shell smooth to a diameter of 16 mm (PI. 5, figs. 20 and 21; PI. 11, fig. 1); first ornament to appear is small umbilical bullae (PI. 5, fig. 13; PI. 11, figs. 2 and 3), which give rise to feeble flexuous prorsiradiate ribs and striae that generally efface on outer flank, although in some specimens there are feeble ribs and depressions on venter (PI. 11, fig. 2). This stage persists to a diameter of 40 mm (PI. 5, figs. 13 and 22; PI. 11, figs. 4 and 5), beyond which strong ventral ribs appear. From this point to maturity (Pis. 1-3; PI. 4, figs. 4-6) there are nine to eleven umbilical bullae per whorl. These arise as low swellings on umbilical wall, strengthen over the shoulder, and are elongate, crescentic, and ofvariable strength within (e.g. PI. 3, fig. 2) and between individuals (compare PI. 1 and PI. 3, figs. 1-3). Where strongly developed they extend as ribs across inner flank before dec1ining (PI. 3, fig. 2); in some cases they appear to subdivide across outer flank (pI. 3, fig. 4) with secondary ribs intercalated (pI. 3, figs. 2 and 4). In other specimens (PI. I, fig. 2; PI. 2, fig. 2; PI. 4, fig. 5) ornament is virtually effaced at mid-flank. In all specimens, ribs strengthen on ventrolateral shoulder, where they are distinctly prorsiradiate, blunt, and rounded, and separated by slightly wider interspaces. All ribs pass straight across venter, and are interrupted by narrow groove marking siphonalline. There are approximately eighty ventral ribs corresponding to the nine to eleven umbilical bullae.
Suture line deeply and intricately subdivided, with symmetrically bifid saddles and markedly retracted
umbilical lobe (text-fig. 4c).
Ward & Kennedy (1993):
Biscay specimens are up to 150 mm diameter.
Coiling moderately involute, with small shallow umbilicus comprising
around 25 percent ofdiameter. Low umbilical wall seems
to have been flattened, with narrowly rounded umbilical shoulder;
whorl section cannot be reconstructed. Juveniles smooth
to around 16 mm. First ornament to appear is small umbilical
bullae, followed by feeble ventral ribs. Strong ventral ribs appear
at about 40 mm diameter. Generally 9-1 1 umbilical bullae per
whorl. These arise on umbilical wall and strengthen markedly
over umbilical shoulder, concave and prorsiradiate on inner
flank, declining around mid-flank, where some appear to subdivide,
although most efface. Strong ribs reappear on ventrolateral
shoulder and strengthen across venter, which they cross
transversely and are coarse, blunt and rounded, numbering
around 80 per whorl.
|Discussion / Comments:
Cobban & Kennedy (1995):
Two specimens, USNM 463 106, 463 107.
The fragmentary nature of this specimen precludes
confident assignation to P. (P.) gollevillensis (see revision
in Kennedy, 1986d), but the high number of intercalated ribs
versus primary ribs, plus effacement of ornament at mid-flank
strongly suggest it belongs to this upper Maastrichtian species.
A second worn fragment, USNM 463106 may also belong here; it has a maximum preserved whorl height of 33.5 mm, a whorl
breadth to height ratio of 0.72, distant umbilical bullae, and
ribs effaced at mid-flank. Presence of umbilical bullae, plus the
numerous short ribs on the outer flank and venter separate the
present material from other Pachydiscus (Pachydiscus) in the
Prairie Bluff Chalk.
Prairie Bluff Chalk at localities 14 and 23.
Discussion. The neotype is a specimen according weIl with the figures of Seunes (1891) and de Grossouvre (1894). It is generally comparable with d'Orbigny's pl. 101, which most authors have used as a basis for the species, whereas d'Orbigny's pI. 102, figs. land 2 are obviously a juvenile Pachydiscus jacquoti Seunes, 1890a.
P. valognensis (Spath, 1922a), the holotype of which is figured for the first time as Plate 2, is inseparable from P. gollevillensis. P. neubergicus nowaki Mikhailov, 1951 (p. 65) is based on the original ofN owak (1913, p. 354, pl. 41, fig. 13); it has a somewhat coarser ventral ornament, but I can see no criteria on which to separate it specifically from P. gollevillensis. Blaszkiewicz (1980, p. 45, pI. 35, figs. 1-3,9) differentiates it from P. gollevillensis sensu stricto on the basis of the more numerous umbilical bullae, wider umbilicus, and more conspicuously reduced costulation in the central parts of the sides. The original ofhis pI. 35, figs. land 9 bearslittle resemblance to the gollevillensis group, however. P. g. armenicus Atabekian and Akopian, 1969 (p. 8, pI. I, fig. 2; pl. 3, figs. land 2) from the Maastrichtian of Armenia has many more umbilical bullae (sixteen to twenty) than the present material (nine to eleven) and sixty-two to seventy-three ventral ribs. It seems distinct enough from the material described here.
P. gollevillensis is a elose ally and, I believe, a descendant ofthe older P. neubergicus (Hauer, 1858) (p. 12 (pars), pl. 2 (3), figs. 1-3 only), the lectotype of which is illustrated in Plate 4, fig. 3. This species has 50% more umbilical bullae than P. gollevillensis, primary ribs that extend across the flank and bi- or trifurcate on the ventrolateral shoulder, far fewer ventral ribs, and a wider umbilicus. Evolution thus involved a reduction in the number of umbilical bullae, effacement of lateral ornament, and an increase in number and decrease in strength of ventral ribs. It is thus perfectly possible that P. g. armenicus may be an intermediate stage in this sequence which has attained the finer ventral ornament of gollevillensis while retaining the style of umbilical ribbing of neubergicus.
Occurrence. Calcaire a Baculites, Upper Maastrichtian of Fresville, Golleville, 'Valognes' and Port Filiolet, Picauville, Manche, France; Hommes Morts, Lleida, Spain; Poland, north Germany, Austria, Armenian SSR, northern Caucasus, Crimea, Bulgaria, the Bithynian Peninsula, Turkey, and Madagascar.
Ward & Kennedy (1993):
Neotype, designated by Kennedy, 1986c, p. 29, is
BMNH C38 179, from the upper Maastrichtian Calcaire a Baculites
of Fresville, Manche, France.
An undeformed topotype is shown in Figure
32.1-32.3 for comparison with the crushed Biscay material.
Pachydiscus valognensis (Spath, 1922) is a synonym of P. (P.)
gollevillensis; the holotype was illustrated by Kennedy (1 986c,
Pl. 2). Pachydiscus (P.) neubergicus nowaki Mikhailov, 1951 (p. 65), based on the original of Nowak (19 13, p. 354, Pl. 41, fig.
13) is also regarded as a synonym, in spite of the views of
Blaszkiewicz (1980, p. 45). Pachydiscus (P.) gollevillensis armenicus
Atabekian and Akopian, 1969 (p. 8, Pl. 1, fig. 2, Pl. 3,
figs. 1, 2; see Figure 29.1-29.6) is here afforded full specific
status. It has many more umbilical bullae (16-20 vs. 9-1 I) and
fewer ventral ribs (62-73 vs. 80). Equally distinctive is P. (P.)
neubergicus neubergicus (Hauer, 1858), described above. It has
14-1 7 umbilical bullae, primary ribs that extend across the flanks
and bi- or trifurcate, far fewer ventral ribs (48-60 vs. 80), and
a wider umbilicus. Pachydiscus (P.) jacquoti jacquoti Seunes,
1890a, also described above, is evolute, with few, distant ribs.
In the Biscay sections this taxon ranges from
the upper half of Member I (lower Maastrichtian) to the top of
Member IV (upper Maastrichtian). Where well dated elsewhere,
the species is restricted to the upper Maastrichtian, with records
from the Calcaire a Baculites of the Cotentin Peninsula, Manche,
France, Hommes Morts, Lleida, Spain, Kunrade and Geulhem,
Limburg, the Netherlands, Poland, north Germany, Austria, the
Armenian SSR, northern Caucasus, Crimea, Bulgaria, Bithynian
Peninsula, Turkey, and Madagascar.
|Cobban & Kennedy (1995):
1853 Ammonites gollevillensis d'Orbigny. - Sharpe (non d'Orbigny, 1850)
: p.48 pl. 17, fig. 2 (= Pachydiscus sharpei Spath)
Ward & Kennedy (1993):
1898 Pachydiscus gollevillensis d'Orbigny. - Kossmat (non d'Orbigny, 1850)
: 97 (162) pl. 15 (21), fig. 1 (=Pachydiscus compressus Spath)
|Was used in synonym list of:
|Stratigraphy - relative ages:
|upper Maastrichtian: Kennedy (1986)
upper Maastrichtian - lower Maastrichtian: ()
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