Pachydiscus jacquoti jacquoti Seunes 1890 from: Ward, P.D..Kennedy, W.J.. (1993): Maastrichtian Ammonites from the Biscay Region (France, Spain) . Memoir (The Paleontological Society), Journal of Paleontology34 (Supplement to Vol. 67)(5) p. 1-58 . |
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Subspecies Pachydiscus jacquoti jacquoti Seunes 1890 |
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Alternative name: |
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Diagnosis / Definition: |
Ward & Kennedy (1993):
Description. -
Biscay specimens range up to 150 mm in diameter,
and are commonest pachydiscid in fauna. Coiling moderately
evolute, with around 40 percent of previous whorl covered.
Whorls expand slowly; umbilicus comprises around 30
percent of diameter. Juvenile shell smooth as can be seen from
inner whorls of large specimens, with ornament first appearing
at estimated diameter of 15 mm. Distant bullae, around eight
per whorl, are first ornament to appear, but by diameter of 80-
90 mm there are up to 14 bullae that give rise to narrow, distant,
straight prorsiradiate ribs that weaken markedly on outer flank,
flex forwards and cross venter in broad convexity. Ribs occasionally
branch in pairs from bullae, or intercalate so that there
are just over twice as many ventral ribs as umbilical bullae.
Beyond 90 mm, bullae may decline as do secondary ribs, leaving
ornament of primary ribs only, weakened markedly over venter. |
Discussion / Comments: |
Ward & Kennedy (1993):
Types. -
Seunes figured three specimens (1890a, PI. 3(2), figs.
1-3) and mentioned others, from the Calcaire a Baculites of the
Cotentin Peninsula, Manche, France, Pyrénées-Atlantiques and
NW of Alcoy, Spain. The original of his Pl. 3(2), fig. 1 should
be designated lectotype, when found (see discussion in Kennedy,
1986c, p. 36).
Discussion.-
Coiling, whorl section (when uncrushed), and
distant, sparse ribbing render Pachydiscus jacquoti jacquoti easily distinguishable from all of the other pachydiscids in the
Biscay fauna; differences are discussed in detail by Kennedy
(1986c, p. 36). Pachydiscus (P.) llareni Wiedmann, 1960 (p. 764,
Pl. 4, fig. 6, Pl. 5, fig. 4) is no more than a crushed P. (P.) jacquoti
jacquoti in our view. Pachydiscus jacquoti australis Henderson
and McNamara, 1985 (p. 76, Pl. 8, figs. 1, 2, 7-10, text-figs.
12a, 13b, 14, 15a) differs from the nominate subspecies in retaining
secondary ribs to maturity, whereas they are lost in the
former. Pachydiscus (P.) jacquoti chilensis Stinnesbeck, 1986 (p.
2 18, Pl. 13, fig. 8, Pl. 1 5, figs. l,3, text-fig. 28a) was differentiated
from the nominate subspecies on the basis of possessing 22-26
ribs per whorl. The holotype is a massive shell, distinguished
by its coarse ribbing, unlike that of the present material.
Occurrence.-
In the Biscay sections this taxon ranges from
near the top of Member I to the top of Member V. Elsewhere,
the nominate subspecies is restricted to the upper Maastrichtian
with records from the area between Gan and Rébénacq, Pyrénées-
Atlantiques, Tercis, Landes, the Calcaire a Baculites of the
Cotentin Peninsula, Manche, all in France; Kunrade, the Netherlands,
Armenia, and Madagascar. |
Systematics: |
38 Ordo Ammonoidea
Subordo Ammonitina
Superfamilia Desmocerataceae
Familia Pachydiscidae
Genus Pachydiscus
Subgenus Pachydiscus
Subspecies Pachydiscus jacquoti jacquoti
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Synonym list: |
Ward & Kennedy (1993):
1987 Pachydiscus (Pachydiscus) jacquoti Seunes. - Kennedy : p.171 pl. 11, figs. 1-4;
pl. 12, figs. 1-3, 8-10;
pl. 13, fig. 1-5;
pl. 14, fig. 4-6, 8-10;
pl. 15, fig. 1-3, 12, 13;
text-fig. 7b
1993 Pachydiscus jacquoti jacquoti Seunes. - Ward & Kennedy : p. 27, 35, 36, 37, 4 fig. 25.8, 25.13, 29.1-29.4, 30.1-30.3, 31.2, 31.3, 31.6, 35.7, 39.4, 40.2
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Stratigraphy - relative ages: |
lower Maastrichtian - upper Maastrichtian: Ward & Kennedy (1993)
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References: |
Seunes,J.. (1890): Recherches géologiques sur les terrains secondaires et l' Eocene inferieur de la region sous-pyrénéene du sud-ouest de la France (Basses-Pyrenees et Landes). p. 250 pp.
Wiedmann,J.. (1960): Le Crétacé supérieur de l'Espagne et du Portugal et ses céphalopodes . C. R. du Congrés des Sociétés Savantes-Dijon, 1959; Colloque sur le Crétacé supérieur francais p. 709-764
Wiedmann,J.. (1964): Le Crétacé supérieur de l'Espagne et du Portugal et ses céphalopodes . Estudios Geologicos Instituto Lucas Mallada Vol. 20 p. 107-148
Kennedy,J.W.. (1986): The Campanian-Maastrichtian ammonite sequence in the environs of Maastricht (Limburg, the Netherlands) . Newsletters on Stratigraphy Vol. 16 p. 149-168
Kennedy,W.J.. (1986): The ammonite fauna of the Calcaire à Baculites (Upper Maastrichtian) of the Cotentin Peninsula (Manche, France) . Palaeontology Vol. 29(1) p. 25-83
Kennedy,W.J.. (1987): The ammonite faunas of the type Maastrichtian, with a revision of Ammonites colligatus Binkhorst, 1861 . Bulletin de l'Institut Royal des Sciences Naturelles de Belgique, Sciences de la Terre Vol. 56 p. 151-267
Ward,P.D.. and Kennedy,W.J.. (1993): Maastrichtian Ammonites from the Biscay Region (France, Spain) . Memoir (The Paleontological Society), Journal of Paleontology34 (Supplement to Vol. 67)(5) p. 1-58
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