Glyptoxoceras rugatum Forbes 1846 from: Ward, P.D..Kennedy, W.J.. (1993): Maastrichtian Ammonites from the Biscay Region (France, Spain) . Memoir (The Paleontological Society), Journal of Paleontology34 (Supplement to Vol. 67)(5) p. 1-58
|Notice: This catalogue page may contain unedited data.
Species Glyptoxoceras rugatum Forbes 1846
|Diagnosis / Definition:
Kennedy & Henderson (1992):
The early growth stages are shown by a series of evenly ribbed and occasionally constricted fragments down to a whorl heights of as little as 3mm. They vary from compressed to subcircular in cross section and are slightly curved, defining obtuse or acute angels, or straight. The same shapes characterize the largest fragments, indicating a planispiral shell, polygonal in outline, and made up of straight shafts connected by obtuse or acute angels. The lectotype of Hamites rugatus (Forbes, 1846, pl. 11, fig. 2) is a wholly septate fragment consisting of two diverging shafts linked by a curved sector (Pl. 1, figs. 1-2). It has a maximum preserved whorl height of 198 mm, and a whorl breadth to height ratio of 0.87; the rib index is 8, the ribs blunt, somewhat weakened on the dorsum but strengthening progressively across the flanks, and strong and transverse on the venter. They are prorsiradiate at the smallest and largest diameters preserved, but markedly rursiradiate on the curved sector. The lectotype of Hamites subcompressus Forbes, 1846 (Pl. 1, figs. 13-16) is a curved body chamber, 150 mm long. The whorl section is compressed oval, with flattened subparallel flanks and a broadly rounded dorsum and venter. The costal whorl breadth to height ratio is 0.73, and the rib index varies between 7 and 8. The ribs are relatively weak and transverse on the dorsum, but strenghten across the flanks, where they are feebly prorsiradiate, and on the venter, where they are transverse. The ribs are distinctly narrower than the interspaces, but show some variation, being blunter on the internal mould than where shell is preserved. The holotype of Glyptoxoceras circulare Shimizu, 1935 (Text-figs. 10-11; Pl. 4, fig. 12) is a broken, straight shaft 95 mm long, with a circular cross section, embedded in matrix. The style of ribbing is, so far as visible, like that of the lectotype of subcompressus, with a rib index of six. Suture (Text-fig. 1A, E) with moderately incised, bifid lobes and saddles.
Ward & Kennedy (1993):
Specimens are crushed fragments, either straight shafts or shafts associated with curved sections. Greatest preserved
whorl height 18 mm. Rib index 6-7. Ribs weak on dorsum,
strengthening over dorsolateral area and flank to reach
maximum development over venter. They are prorsiradiate,
straight, and narrower than interspaces. Sutures not seen.
|Discussion / Comments:
Kennedy & Henderson (1992):
Lectotype, here designated, is BMNH C51110, the original of Forbes, 1846 (pl. 11, fig. 2), GSC R10499.
The lectotype, here designated, of Hamites subcompressus Forbes, 1846, is shown in Plate 1, figures 12-16. it is broken into three parts (Pl. 1, fig. 16). The middle fragment (A) as noted by Philipps (1977, p. 125), GSC R10491, is the original of Kossmat (1895, pl. 19(5), fig. 10), and possibly the original of Forbes (1846, pl. 11, fig. 6). The adapical fragment (B) corresponds with Forbe's plate 11, figure 4a, c, the figured syntype of Hamites indicus. Fragment C was never figured. The lectotype, here designated, of Hamites nereis Forbes, 1846, plate 10, figure 7, is BMNH C51109, GSC R10502. In each case lectotypes are designated because Forbes cited the 'dimension of largest specimen', indicating that he possessed more than one of each species. The holotype of Glyptoxoceras circulare Shimizu, 1935 (text-figs. 10-11) is BMNH C51112. Possible paralectotypes of Hamites subcompressus are BMNH C51112, the original of Kossmat (1895, pl. 19(5), fig. 4c = GSC R10497; Pl. 2, fig. 16), and BMNH C 51103, the original of Kossmat (1985, pl. 19(5), fig. 11; Pl. 1, fig. 9, herein). Topotypes BMNH C51125 (Pl. 1, fig. 5), C51131 and C51139 are possible syntypes of Hamites indicus that belong to the present species, as are BMNH C51120-51122 (Pl. 1, figs. 10-11; Pl. 4, fig. 2), syntypes of Hamites undulatus Forbes, 1846; and BMNH C51096-51099, which are syntypes of Hamites largesulcatus Forbes, 1846 (Pl. 3, figs. 1-3; Pl. 4, figs. 13-15)(all ex Kaye and Cunliffe Collection). Other topotypes are BMNH C4049 (5 fragments, no history), C4050 (12 fragments, no history); C4109 (history uncertain, the original of Shimizu, 1935, figs. 6-9), C5114-5116 (no history), BMNH C24201 (4 fragments, ex Kaye Collection), C51126-51129 (ex Geological Society Collection), and C2402 (ex Kaye Collection). All are from the Valudavur Formation of Pondicherry, South India.
We regard Hamites rugatus, H. nereis and H. subcompressus of Forbes (1846) as synonyms, and as first revising authors select the name rugatus for the species. Also regarded as conspecific is G. circulare, and some of the syntypes of Hamites indicus of Forbes. The characteristic features of the species are the polygonal coiling, relatively large size, and rib density. When compared with other species in the Ponicherry fauna. Glyptoxoceras tenuisulcatum (Forbes, 1846) has an initial helix, and much finer ribbing that is oblique on the venter of the shaft, rather than being transverse (Pl. 2, figs. 2, 6, 8, 30). Glyptoxoceras largesulcatum (Forbes, 1846)(Pl. 3, figs. 4-9; Pl. 4, figs. 16-18) has very distant annular ribs, and is only known as straight shafts. G. indicum (Forbes, 1846) (Pl. 1, figs. 1, 3-5, 7, 9, 12-13) has a quite different coiling, with an initial helix, and oval planispiral later whorls, being adult at much smaller size (Pl. 1, figs. 3-4).
When these species are removed from the Pondicherry fauna, the numerous remaining fragments from a species that is variable in whorl section and ribbing density, but of similar coiling and ribbing style. Hamites nereis (Forbes, 1846) (Pl. 2, figs. 10-11), referred to Pseudoxybeloceras by some authors, is a pathological rugatum in which ventral ribbing is interrupted, as a result of non-lethal injury to the mantle margin (Pl. 2, fig. 10).
Of species from Pondicherry illustrated by d'Orbigny (1847a), one of the specimens of Hamites acuticostatus (pl. 3, figs. 11, 12) may belong here; it is shown as being 40 mm long, with a compressed whorl section, and rib index of 4. The second specimen illustrated by d'Orbigny (pl. 3, figs. 9-10) seems to be a G. largesulcatum. The Hamites simplex of d'Orbigny from Pondicherry (pl. 3, figs. 15-16) is a curved fragment 30 mm long, and is clearly G. rugatum.
We refer a number of non-Indian records to G. rugatum on the basis of coiling style, ribbing and age (see synonymy), but reject small fragments that are significantly older where the coiling style is unclear, believing them to be unidentifiable as to genus in some cases. Thus fragments from the Upper Maastrichtian of the Maastricht area, described as Hamites rotundus by Binkhorst (1861) are referred to G. rugatum (see also Kennedy, 1987), as is G. recticostatus (Seunes, 1890) from the Upper Maastrichtian of southeastern France, and G. braziliense (Maury, 1930), from Brazil. Study of more than 100 Glyptoxoceras from the Upper Maastrichtian Miria Formation of western Australia (to be described elsewhere), has convinced us that G. indicum, G. circulare, G. nipponicum, and G. bullarense, Neohamites giralensis, N. rugatus, N. cardabiensis, N. largesulcatus, and N. soufoulisi of Brunnschweiler (1966) represent but a single form, conspecific with Glyptoxoceras rugatum.
The Glyptoxoceras subcompressum of Ward (1976, p. 456, pl. 1, figs. 1-5; text-fig. 3) and Ward and Westermann (1976, text-figs. 1-3) has a quite different, ellipsoidal coiling when compared to the Pondicherry material, with an initial helix with its axis of coiling at 90° to that of the planispiral whorls; it is of late Santonian or early Campanian age.
Maastrichtian of south India, Brazil, Chile (?), Western Australia, northern Spain, south-east France, and the Maastricht area in the Netherlands and Belgium.
Ward & Kennedy (1993):
This species is based upon a series of fragments from
the Valudavur Formation of Pondicheny, south India, in the
collections of the Natural History Museum, London, and lectotype
designation is deferred until this material is published
(Kennedy and Henderson, in prep.).
These crushed fragments provide only rib density,
style, and direction as clues to specific identity, but a comparison
with the largest Indian example of Glyptoxoceras rugatum
in the Forbes Collection reveals no obvious differences.
Glyptoxoceras largesulcatum (Forbes, 1 846) (p. 1 1 7, Pl. 11, fig.
1; see Fig. 18.13) is easily differentiated by the low rib index
(4.5) and large size. Glyptoxoceras tenuisulcatum (Forbes, 1846)
(p. 116, Pl. 11, fig. 3) is in contrast densely and finely ribbed,
with a circular cross section. Glvptoxoceras indicum (Forbes,
1846) p. 1 16, Pl. 1 1, fig. 4) (see Matsumoto, 1959, p. 167, Pl.
4 1, figs. 2-6, text-fig. 80) has a different coiling mode, seemingly
lacking the long straight to slightly curved shafts of G. rugatum.
with a circular whorl section.
In the Biscay sections this taxon ranges from
the upper part of Member I (lower Maastrichtian) to the upper
half of Member IV (upper Maastrichtian). The taxon shows a
discontinuous range across the basin; at Zumaya it is one of the
commonest ammonites, whereas it is rare or absent at the other
Biscay sections. Immel, Klinger, and Wiedmann (1982) referred
fragments from as low as the lower Santonian to the present
species, but it is perhaps questionable how many of these generalized
diplomoceratid scraps are conspecific. The most convincing
records are Maastrichtian. The recorded geographic range
from this stage is southeastern France, Spain, the Netherlands,
south India, and western Australia.
Species Glyptoxoceras rugatum
Species Glyptoxoceras rugatum
|Kennedy & Henderson (1992):
1847 Hamites simplex d'Orbigny. - d'Orbigny
: pl. 3; fig. 15-17 (non d'Orbigny, 1842)
1861 Hamites rotundus Sowerby?. - Binkhorst
: p.34 pl. 5b, fig. 2-4;
pl. 5c, fig. 1
Ward & Kennedy (1993):
1992 Glyptoxoceras rugatum Forbes. - Kennedy & Henderson
: p. 697, 701, 705, 71 pl. 1, fig. 1-2, 5-16;
pl. 2, fig. 10-11, 14-29;
pl. 3, fig. 1-3;
pl. 4, fig. 2, 12-15;
text-fig. 1A, E
|Stratigraphy - relative ages:
|upper Maastrichtian - lower Maastrichtian: Ward & Kennedy (1993)
Maastrichtian: Kennedy & Henderson (1992)
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