Eubaculites carinatus Morton 1834 from: Ward, P.D..Kennedy, W.J.. (1993): Maastrichtian Ammonites from the Biscay Region (France, Spain) . Memoir (The Paleontological Society), Journal of Paleontology34 (Supplement to Vol. 67)(5) p. 1-58 | . |
Notice: This catalogue page may contain unedited data.
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Species Eubaculites carinatus Morton 1834 |
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| Diagnosis / Definition: |
Cobban & Kennedy (1995):
DESCRIPTION.-
Holotype, a phosphatic internal mold of a body
chamber and parts of two camerae, is 47.5 mm long with whorl
breadth to height ratio 0.74 and angle of taper of 7 degrees.
Whorl section pyriform with flattened dorsum, narrowly rounded
dorsolateral margin, and broadly rounded flanks; greatest
breadth below mid-flank. Outer flanks convergent. A faint longitudinal
groove separates flanks from narrow tabulate venter
with very narrowly rounded ventrolateral shoulder. Rib index
2.5. Ribs broad, strong and concave, as wide as interspaces,
extending from dorsolateral margin to middle third of flank. On
outer third ribs weaken, break down into riblets and striae, and
project strongly forward to ventrolateral margin, where they flex
back and are feebly convex. They strengthen markedly on shoulder
and are coarse and distant on venter, which is conspicuously
serrated in profile; ventral ribs approximately twice as numerous
as lateral. Ribs weaken markedly over dorsum and reduce to
mere low convex folds. Numerous fragments match the type
and have whorl heights of up to 19 mm. Variation consists
chiefly of differences in strength of flank ornament and density
with a rib index of 2.5-3. Venters vary from strongly ribbed
and serrate to smooth; dorsums from smooth to delicately ribbed
and striate. Somewhat different, but also regarded as intraspecific
variations, are weaker ribbed individuals with rib index of
up to 4, and a more compressed whorl section (down to 0.6);
these in turn connect to nearly smooth variants and those with
growth lines and striae only on flank. Suture moderately incised
with broad bifid saddles, narrow L, and broad U.
Kennedy & Cobban (1996):
DESCRIPTION.-
Specimens are phosphatized internal molds of
body chamber. Whorl section compressed, dorsum broad, feebly
convex, dorsal flanks convex, converging ventrally to narrow,
tabulate venter. Flanks ornamented by coarse crescentic ribs on
dorsal two-thirds of flank, sweeping forward and declining on
ventral third of flank, where additional riblets and striae may
develop (Figure 3.1 1); rib index 1.5-2. Venter ornamented by
coarse transverse riblets, 5 or 6 in distance equal to whorl height
(Figure 3.3, 3.9, 3.10).
Ward & Kennedy (1993):
Description. -
Specimens crushed. Shell straight, and expanding
slowly. Largest specimen seen has whorl height of 18 mm.
Flank ornament of coarse concave crescentic ribs, 2.5-4.0 in a
distance equal to whorl height. They are strongest on dorsolateral
area, and extend across most of flank, projecting forward
and declining toward ventrolateral margin, which may be flanked
by shallow groove. Ventrolateral shoulders sharp, venter tabulate.
Landman et al. (2004):
DESCRIPTION.-
The collection includes both
robust and slender forms, presumably macroconchs
and microconchs, respectively. These
differences are expressed in the angle of taper,
although this is also very dependent on
ontogenetic stage (table 6). The dorsal onehalf
of the flanks are covered with broad,
asymmetrically crescentic ribs with a rib index
of 2–3. The narrow tabulate venter is
serrated with a rib index of 6–8. We describe and illustrate 15 specimens to convey the
range of variation of our material.
AMNH 47418 is a piece of body chamber
49.7 mm long with whorl heights of 15.1 and
17.1 mm at the adapical and adoral ends, respectively
(fig. 27F–H). The angle of taper
is 3.28. The whorl section at the adoral end
is ovoid with maximum width at one-third
whorl height both costally and intercostally;
the intercostal ratio of whorl width to height
is 0.60. The dorsum is very broadly rounded
and the dorsolateral shoulder is fairly abruptly
rounded. The flanks are broadly rounded
and converge to the venter. The narrow, flat
venter is bordered by a faint longitudinal
groove.
The dorsum shows low, convex folds
spaced at distances of approximately 5 mm
near the adoral end. There are four large,
slightly crescentic nodate swellings on the
inner two-thirds of the flanks. The swellings
become progressively more widely spaced
adorally; the distance between the two most
adoral swellings is approximately 14 mm;
the rib index is approximately 2. The nodes
are strongest just ventral of the dorsolateral
margin. The venter is covered by coarse,
swollen ribs that show a slight forward projection.
They gradually become more widely
spaced adorally so that the distance between
rib crests at the adoral end of the specimen
is approximately 4 mm; the rib index is 6.
AMNH 47506 is a body chamber fragment
56.9 mm long with whorl heights of 11.7 mm
and 13.4 mm at the adapical and adoral ends,
respectively (fig. 27Q–T). The angle of taper
is 3.08. The ratio of whorl width to height at
the adoral end is 0.70. There are low folds
on the dorsum. The inner two-thirds of the
flanks bear strong crescentic ribs spaced at
distances of approximately 10 mm at the adoral
end; the rib index is 2.5. The narrow,
tabulate venter is ornamented with strong,
transverse ribs with a slight adoral projection,
producing a serrated appearance; the rib
index is 5.
AMNH 47409 is a large fragment of a
body chamber 58.2 mm long with whorl
heights of 16.7 mm and 19.0 mm at the adapical
and adoral ends, respectively (fig. 28H–
K). The ratio of whorl width to height at the
adoral end is 0.72. The whorl section is
ovoid with maximum width at one-third whorl height. The dorsum is flat and the dorsolateral
margin is fairly sharply rounded.
The inner flanks are broadly rounded and the
outer flanks converge toward the venter. The
venter is set off by a weak, longitudinal furrow
and the ventrolateral shoulder is fairly
abruptly rounded; the venter is narrow and
flat.
Weak swellings occur on the dorsal onehalf
of the flanks with a distance of 13.5 mm
between the two most adoral swellings; the
rib index is approximately 2.5. The venter is
covered with swollen ribs that show a slight
forward projection. The ribs are spaced at
distances of 4 mm so that the rib index is
approximately 6.5.
AMNH 47412 is the adapical part of a
body chamber 48.4 mm long (fig. 28L–P).
The whorl heights are 14.1 mm and 15.7 mm
at the adapical and adoral ends, respectively.
The whorl section at the adoral end is pyriform
with an intercostal ratio of whorl width
to height of 0.73. The angle of taper is 2.28.
The dorsum is nearly flat to very broadly
rounded and the dorsolateral margin is fairly
abruptly rounded. The flanks are broadly
rounded with maximum width at one-third
whorl height. The outer flanks converge to
the venter. The venter is flat and narrow and
is bordered by a ventrolateral furrow producing
a keel. The ventrolateral shoulder is
abruptly rounded.
There are very faint, convex folds on the
dorsum. Broad, crescentic ribs cover the dorsal
one-half of the flanks. On the left side,
these ribs are of variable strength and spacing
with a rib index of 4, whereas on the
right side, these ribs are more nodate with a
rib index of 2. Ribs slant strongly forward to
the ventrolateral margin. The venter is covered
with fairly coarse ribs, which show a
weak forward projection, producing a serrated
appearance; the rib index is 8.
AMNH 47416 is a fragment of a body
chamber 45.6 mm long with whorl heights
of 13.9 mm and 17.5 mm at the adapical and
adoral ends, respectively (fig. 28Q–T). The
ratio of whorl width to height at the adoral
end is 0.69. Broad, weak crescentic swellings
cover the dorsal one-half of the flanks with
a rib index of approximately 3. The left side
shows a dent near the adapical end, which is
probably the result of an injury. The venter is mostly eroded away but shows three swollen
transverse ribs on the adoral end. The ribs
are spaced at distances of 2 mm so that the
rib index is approximately 10.
MAPS A2058a4 is a strongly ribbed fragment
of a body chamber 59.7 mm long with
whorl heights of 15.6 mm and 16.4 mm at
the adapical and adoral ends, respectively
(fig. 29A–E). The intercostal ratio of whorl
width to height at the adoral end is 0.70. The
angle of taper is 1.28. There is a shallow,
longitudinal groove on either side of a fastigiate
to tabulate venter. The dorsum is covered
with low folds that show a relatively
strong adoral projection. There are broad,
asymmetrically crescentic ribs on the dorsal
one-half of the flanks with a rib index of 3.
The venter is notched with swollen ribs
showing a slight adoral projection; the rib index
is 5.5.
AMNH 47410 is a body chamber fragment
with some iridescent shell on the adapical
end; the angle of taper is 3.88 (fig. 29F–I). It
is 46.5 mm long with whorl heights of 13.4
mm and 15.7 mm at the adapical and adoral
ends, respectively. The intercostal ratio of
whorl width to height at the adoral end is
0.73. The whorl section is pyriform with a
broadly rounded to flat dorsum and fairly
sharply rounded dorsolateral margin. The inner
flanks are broadly rounded and the outer
flanks converge to the venter. The venter is
bordered by a faint longitudinal groove. The
ventrolateral shoulder is fairly steeply rounded
and the venter is narrow and tabulate.
Swollen, concave ribs cover the inner onehalf
of the flanks. The distance between rib
crests is approximately 8 mm near the adoral
end so that the rib index is approximately 3.
Ribs weaken and break down into riblets on
the outer one-third of the flanks. They slant
strongly forward at an angle of approximately
208. The ribs strengthen on the venter and are spaced every 2 mm near the adoral end,
so that the rib index is approximately 7.
AMNH 47511 is a coarsely ornamented
fragment of the adapical part of a body
chamber 38.8 mm long with whorl heights
of 12.5 mm and 15.0 mm at the adapical and
adoral ends, respectively (fig. 29J–M). The
intercostal ratio of whorl width to height at
the adoral end is 0.72. The angle of taper is
4.28. There are low folds on the dorsum. The
inner two-thirds of the flanks bear strong,
crescentic ribs spaced at distances of approximately
10 mm at the adoral end, so that the
rib index is 2.5. The venter is narrow and
tabulate and ornamented with strong, transverse
ribs that show a slight adoral projection,
producing a serrated appearance; the rib
index is 5.
AMNH 47415 is a small specimen of a
body chamber 35.6 mm long, with relatively
coarse ornament (fig. 29N–R). The whorl
heights are 12.3 mm and 14.1 mm at the
adapical and adoral ends, respectively. The
intercostal ratio of whorl width to height at
the adoral end is 0.75. The angle of taper is
3.68. The inner one-half of the flanks are covered
with four slightly crescentic nodelike
swellings with the interspaces between them
as broad as the swellings themselves. The
distance between node crests is 9 mm, so that
the rib index is approximately 2.5. The ribs
slant strongly forward on the outer one-half
of the flanks at an angle of approximately
208. The narrow tabulate venter is covered
with swollen, convex ribs spaced at distances
of 2.5–3.0 mm, yielding a rib index of 6.
AMNH 47413 is a small piece of the adoral
part of a phragmocone and the adapical
part of a body chamber of a specimen 35.9
mm long (fig. 29V–X). The whorl heights at
the adapical and adoral ends are 9.2 mm and
11.9 mm, respectively. The ratio of whorl
width to height at the adoral end is 0.75. The angle of taper is 5.28. The inner one-half of
the flanks are covered with broad, crescentic
ribs with the interspaces between them as
broad as the ribs themselves. The distance
between rib crests at the adoral end of the
specimen is 6 mm; the rib index is approximately
3. The narrow tabulate venter is
smooth.
AMNH 47504 is a fragment of a body
chamber 35.3 mm long with whorl heights
of 9.7 mm and 11.0 mm at the adapical and
adoral ends, respectively (fig. 29Y–b). The
ratio of whorl width to height at the adoral
end is 0.79. The angle of taper is 4.08. The
dorsal one-half of the flanks bear nodelike
swellings with a rib index of approximately
2. The ribs weaken and slant strongly forward
on the outer one-half of the flanks.
They are transverse on the venter, with a rib
index of 6.
AMNH 47270 (not illustrated) is a body
chamber fragment 55.8 mm long with a
whorl height of 22.6 mm at the adoral end.
The whorl section is compressed ovoid at the
adoral end with a ratio of whorl width to
height of 0.75. The dorsum is broad and
nearly flat and the dorsolateral shoulder is
fairly abruptly rounded. The inner flanks are
broadly rounded and nearly parallel and the
outer flanks converge toward the venter. The
venter is narrow and tabulate and the ventrolateral
shoulder is abruptly rounded. There is
a shallow longitudinal groove that borders
the venter. The dorsum is covered with low,
convex folds. The venter is nearly smooth
and the dorsal one-half of the flanks bear
widely spaced nodelike swellings with a rib
index of approximately 2.5.
AMNH 47444 is one of the smallest fragments
in our collection, a piece of phragmocone,
13.6 mm in length, with whorl
heights of 9.9 mm and 10.6 mm at the adapical
and adoral ends, respectively (not illustrated).
The ratio of whorl width to height at
the adoral end is 0.74. There are two nodelike
swellings on the inner flanks with a rib index of 2. The ribs are very weak on the
tabulate venter with a rib index of 4.5 based
on a distance of 3 mm between the two most
adoral ribs.
MAPS A2058a6 (not illustrated) is a completely
septate fragment 63.0 mm long with
whorl heights of 16.0 mm and 18.7 mm at
the adapical and adoral ends, respectively.
The whorl section at the adoral end is compressed
ovoid with a ratio of whorl width to
height of 0.75. The dorsum is very broadly
rounded and the dorsolateral margin is well
rounded. The flanks are broadly rounded and
converge toward the venter with maximum
width at midwhorl height. The venter is bordered
by a shallow, longitudinal depression
and the ventrolateral shoulder is fairly sharply
rounded; the venter is broadly rounded to
flat. The inner one-half of the flanks are
smooth except for weak, nodelike swellings
with a rib index of approximately 2. Broad,
transverse ribs cover the venter with a rib
index of 7. Parts of the last and next to last
suture are illustrated in figure 30B. The sutures
are worn but show a bifid L/U, a broad,
bifid U, and a narrow I.
AMNH 47159 is a fragment of the adapical
end of a body chamber 43.4 mm long
with whorl heights of 11.2 mm and 13.4 mm
at the adapical and adoral ends, respectively
(fig. 33U–Y). The ratio of whorl width to
height at the adoral end is 0.74. The venter is
narrow and nearly flat and is bordered by a
longitudinal furrow. There are weak crescentic
swellings on the dorsal one-half of the flanks
with a rib index of approximately 3. The venter
is ornamented with weak, transverse ribs with
a rib index of approximately 8.
Landman et al. (2007):
DESCRIPTION.-
The apical angle ranges
from 1.9 to 4.5u (table 5). The whorl cross
section is compressed ovoid. The intercostal
ratio of whorl width to height ranges from
0.68 to 0.77 and the costal ratio of whorl
width to height ranges from 0.69 to 0.84
(table 5). These ratios are nearly constant
throughout ontogeny. In a plot of whorl
width versus height in intercostal section, the
y-intercept is approximately 0 (at x 5 0),
indicating that the growth of whorl width
versus height is nearly isometric (fig. 35).
The dorsum is almost flat to very broadly
rounded and the dorsolateral margin is fairly
abruptly rounded. The inner flanks are
broadly rounded with maximum width at
one-third whorl height both costally and
intercostally. The outer flanks converge to
a fastigiate to tabulate venter. The venter is
bordered by a shallow, longitudinal groove
on each side.
The inner one-half of the flanks are
covered with weak to strong, slightly crescentic
nodate swellings with a rib index of
1.2–2.8. The nodes are particularly well
developed in the two largest specimens,
MAPS A2058b1 (fig. 29) and AMNH
50766 (fig. 30A–E), but also occur in very
small specimens such as AMNH 50771
(fig. 32E–I). In other specimens such as
AMNH 50779 (fig. 33N–Q), the nodes are
relatively weak. The venter is nearly smooth
to strongly ornamented with transverse ribs
that cross the venter with a slight adoral
projection, producing a serrated appearance;
the rib index ranges from 3.5 to 8.5 and
averages 6 (table 5).
MAPS A2058b1 (fig. 29) and AMNH
50737 (fig. 31I) preserve a mature modification
at the aperture. In both instances, there is a short dorsal and long ventral rostrum,
each of which tapers gradually to a rounded
tip. It is interesting that these two specimens
are very different in size, with whorl heights
at the aperture of 21.7 and 12.2 mm,
respectivey, suggesting that they may represent
antidimorphs.
One specimen (AMNH 50434), a small
fragment with a whorl height of 16.8 mm at
the adoral end, contains part of a convex
structure inside the adoral end of the body
chamber (fig. 32J–N). The structure looks
suspiciously like part of a lower jaw (aptychus).
It is ornamented with ribs that parallel
the outer margin. However, the ribs are not
as rugose as those on baculite lower jaws
(Landman et al., in press) and, in any event,
not enough of the structure is preserved to
make a positive identification.
The suture shows a bifid E/L, a narrow,
bifid L, a broad, bifid L/U, a bifid U, and
a narrow I (fig. 34). |
| Discussion / Comments: |
Cobban & Kennedy (1995):
TYPE.-
The holotype, by monotypy, is no. 72866 in the collection
of the Academy of Natural Sciences of Philadelphia, the
original of Morton, 1834, plate 13, figure 1, from Prairie Bluff,
Alabama.
MATERIAL.-
Twenty figured specimens (USNM 463 174463 193)
and numerous other specimens.
DISCUSSION.-
Baculites carinatus Morton, 1834 is the earliest
available name for widely occurring Eubaculites with a tabulate
venter and crescentic flank ribs. The species has been described
and discussed at length by Kennedy (1986d, as E. lyelli, and
Henderson et al., 1992), who described the intraspecific variation
shown by the species, and discussed its synonyms, the most
important of which are Baculites lyelli d'orbigny, 1847, based
on material from Quiriquina Island, Chile; Baculites tippaensis
Conrad, 1858 and B. spillmani Conrad, 1858 originally described
from the Owl Creek Formation of Mississippi; plus E.
kossmati Brunnschweiler, 1966, E. compressum Brunnschweiler,
1966, and E. multicostatus Brunnschweiler, 1966, originally
described from Western Australia. Many references to E. ootacodensis
(=E. labyrinthicus herein) actually refer to E. carinatus,
as is clear from the synonymy of E. labyrinthicus given above.
Presence of a pair of flank tubercles rather than crescentic flank
ribs distinguish populations of E. vagina (Forbes, 1846) (see
revision in Kennedy and Henderson, 1992b), although smooth
variants of the two species may be inseparable. Populations of
E. latecarinatus (Brunnschweiler, 1966) have smooth flanks (see revision in Henderson et al., 1992). Eubaculites simplex (Kossmat,
1897) (see revision in Henderson et al., 1992) are very
compressed, smooth or with feeble flank undulations, and have
a fastigiate to narrowly tabulate venter at a growth stage when
E. carinatus invariably possesses a tabulate venter.
OCCURRENCE.-
Maastrichtian, Prairie Bluff Chalk at localities
2, 5-9, 16, 31, 32, 36, 37, 40 and 44-47.
The species also occurs in the Owl Creek Formation in Missouri,
Mississippi and Tennessee, Corsicana Formation in
northeast Texas, and as remaink fossils at the base of the Hornerstown
Sand in New Jersey. There are also records from The
Netherlands, southwestern France, northwestern Spain, Austria,
Mozambique, Zululand (South Africa), South India, Western
Australia, Argentina, Chile and California.
Kennedy & Cobban (1996):
TYPE.-
The holotype, by monotypy, is no. 72866 in the collection
of the Academy of Natural Sciences of Philadelphia, the
original of Morton, 1834, pl. 13, fig. 1, from Prairie Bluff, Alabama.
DISCUSSION.-
The present specimens are larger than most individuals
of this species from the Owl Creek Formation, Prairie
Bluff Chalk and Corsicana Formation (Cobban and Kennedy,
1995, in press; Kennedy and Cobban, 1993), but find a match
in individuals of comparable size from the Mira Formation of
Western Australia (Henderson et al., 1992).
MATERIAL.-
Three specimens, USNM 1269la-c, ex J. B. Marcou
Collection, labelled 'New Jersey' are, by their preservation,
from the base of the Hornerstown.
OCCURRENCE.-
Maastrichtian base of Hornerstown Formation
in New Jersey; Prairie Bluff Chalk of Mississippi and Alabama;
Owl Creek Formation in Missouri, Mississippi and Tennessee;
Corsicana Formation in northeast Texas. There are also records
from the Netherlands, southwestern France, northwestern Spain,
Austria, Mozambique, Zululand (South Africa), South India,
Western Australia, Argentina, Chile and California.
Ward & Kennedy (1993):
Type.-
Holotype, by monotypy, is No. 72866 in the collections
of the Academy of Natural Sciences, Philadelphia, from
the Maastrichtian Prairie Bluff Chalk of Alabama.
Discussion. -
The closely spaced ribbing, extending across the
greater part of the flank, separates these specimens from other
baculitids in the Biscay fauna, even when the otherwise diagnostic
tabulate Eubaculites venter is invisible. The synonymy
of the species of Eubaculites is reviewed at length by Kennedy
(1987, p. 192) and Henderson et al. (1992), who modified
the conclusions of Klinger (1976, p. 83 et seq.). Eubaculites
ootacodensis (Stoliczka, 1866) (p. 199, PI. 90, figs. 14, ?I 5) has
ribs combined with a fastigiate venter. Eubaculites vagina
(Forbes, 1846) (p. 114, PI. 10, fig. 4) has a tabulate venter, but
is bituberculate. Eubaculites latecarinatus (Brunnschweiler, 1966)
(p. 33, Pl. 3. figs. 13, 14, Pl. 4, figs. 11-14, text-fig. 20) has a
serrated tabulate venter and smooth flanks.
Occurrence.-
In the Biscay sections this taxon is restricted to
Member IV (upper Maastrichtian). Elsewhere, the species occurs
in both lower and upper Maastrichtian, where well dated, with
records from northern Spain, the Pyrénées-Atlantiques and Petites-
Pyrénées, Haute Garonne in France, northern Spain, the
Netherlands, Neuberg, Steiermark, Austria, Zululand, South Africa,
south India, western Australia, Argentina, and Chile.
Landman et al. (2004):
TYPE.-
The holotype, by monotypy, is
ANSP 72866, the original of Morton, 1834:
pl. 13, fig. 1, from the Maastrichtian Prairie
Bluff Chalk of Alabama.
MATERIAL.-
There are 77 specimens, 5 of
which are float, from the upper part of the
New Egypt Formation and basal part of the
Hornerstown Formation, AMNH loc. 3345,
Parkers Creek, northeastern Monmouth
County, New Jersey, and 1 specimen (AMNH 47159) from 1.5–2 m below the
base of the Hornerstown Formation, AMNH
loc. 3346, just upstream from AMNH loc.
3345. All of the specimens are internal molds
of fragments less than 75 mm in length. A
few specimens retain the original shell material
of the septa. There are many more body
chamber pieces than phragmocone pieces—
there are only four completely septate fragments
(5% of the total).
DISCUSSION.-
This species is characterized
by a narrow, tabulate venter ornamented with
ribs producing a serrated appearance, and
broad, asymmetrically crescentic ribs on the
dorsal one-half of the flanks. The full range
of variation of this species has been recently
discussed by Klinger and Kennedy (1993), Cobban and Kennedy (1995), and Kennedy
and Cobban (2000).
OCCURRENCE.-
This species occurs in New
Jersey in the upper part of the New Egypt
Formation and as reworked material at the
base of the Hornerstown Formation, near Eatontown,
northeastern Monmouth County; at
the top of the New Egypt Formation in the
Crosswicks Creek Basin, southwestern Monmouth
County (Landman et al., in prep. b);
and in the New Egypt Formation at the Inversand
Pit, near Sewell, Gloucester County
(Kennedy and Cobban, 1996). It also occurs
at the top of the Tinton Formation and as
reworked material at the base of the Hornerstown
Formation near Freehold, central
Monmouth County (Landman et al., in prep.
a). Kennedy and Cobban (1996: fig. 3.1–3.3,
3.7–3.12) recorded three specimens of this
species (USNM 12691a–c), ex J.B. Marcou
collection, labeled ‘‘New Jersey’’, which
they inferred to be from the base of the Hornerstown
Formation. This species is reported
elsewhere on the Gulf and Atlantic Coastal
Plains from the top of the Corsicana Formation
and as reworked material at the base
of the Kincaid Formation, Falls County, Texas
(Kennedy et al., 2001); the Owl Creek
Formation, Mississippi, Missouri, and Tennessee
(Kennedy and Cobban, 2000); the
Prairie Bluff Chalk, Alabama and Mississippi
(Cobban and Kennedy, 1995); and the
Severn Formation, Maryland (Kennedy et
al., 1997; Landman et al., 2004). It is also
reported from southeast France, Austria, the
Netherlands, Zululand (South Africa), Mozambique,
Madagascar, southern India, Western
Australia, Chile, Argentina, and California.
It ranges from the upper lower to the
upper upper Maastrichtian (Klinger et al.,
2001).
Landman et al. (2007):
TYPE,-
The holotype, by monotypy, is
ANSP 72866, the original of Morton, 1834:
pl. 13, fig. 1, from the Maastrichtian Prairie
Bluff Chalk of Alabama.
MATERIAL.-
Approximately 50 specimens
in the AMNH collections and 55 specimens
in the MAPS collections from the upper part
of the Tinton Formation, mainly the Pinna Layer, and, more rarely, as reworked material
at the base of the Hornerstown Formation,
Manasquan River Basin, central Monmouth
County. The majority of specimens
consist of fragments of body chambers,
nearly all of which are uncrushed. Most
specimens are juveniles and range in length
from 20 to 60 mm (fig. 35), but two specimens,
probably parts of adults, are exceptionally
long with lengths of 117 and
141 mm. Specimens in the Pinna Layer
sometimes occur in clusters (fig. 6E), which
are probably biological in origin.
DISCUSSION.-
The taphonomy of these
specimens evidently favored gross ornament
over fine ornament. Thus, delicate growth
lines are missing but large nodes are well
preserved. The fact that two specimens retain
mature modifications at the aperture suggests
that these animals lived nearby and did not
suffer much postmortem transport. This is
consistent with the observation that many
specimens occur in monospecific clusters that
may be related to group feeding or the result
of nearly simultaneous death after spawning
or mating. With the exception of one specimen
(AMNH 51327) that bears a single thread of
a bryozoan colony, there are no epizoans.
As summarized by Klinger and Kennedy
(2001: 62), the adult size of this species in
North America and Europe is much smaller
than that in Argentina and Zululand. The
maximum whorl height of a specimen from the Manasquan River Basin is 25.4 mm
whereas the maximum whorl heights of
specimens from Argentina and Zululand are
approximately 90 mm.
OCCURRENCE.-
Top of the Tinton Formation,
mainly the Pinna Layer, and, rarely, as
reworked material at the base of the Hornerstown
Formation, Manasquan River Basin,
central Monmouth County, New Jersey.
Elsewhere in New Jersey, this species occurs
in the upper part of the New Egypt Formation
and as reworked material at the base of the Hornerstown Formation, near Eatontown,
northeastern Monmouth County
(Landman et al., 2004b); the top of the New
Egypt Formation in the Crosswicks Creek
Basin, southwestern Monmouth County
(Landman et al., in prep.); and the New
Egypt Formation and as reworked material at
the base of the Hornerstown Formation at the Inversand Pit, near Sewell, Gloucester County
(Kennedy and Cobban, 1996). Kennedy
and Cobban (1996: fig. 3.1–3.3, 3.7–3.12)
recorded three specimens of this species
(USNM 12691a–c), ex J.B. Marcou collection,
labeled ‘‘New Jersey,’’ which they
inferred to be from the base of the Hornerstown
Formation. Elsewhere on the Atlantic
Coastal Plain, this species occurs in the
Severn Formation, Prince Georges County,
Maryland (Kennedy et al., 1997), and Anne
Arundel County, Maryland (Landman et al.,
2004a). On the Gulf Coastal Plain, this species
is reported from the top of the Corsicana
Formation and as reworked material at the
base of the Kincaid Formation, Falls County,
Texas (Kennedy et al., 2001); the Owl Creek
Formation, Mississippi, Missouri, and Tennessee
(Kennedy and Cobban, 2000); and the
Prairie Bluff Chalk, Alabama and Mississippi
(Cobban and Kennedy, 1995). Klinger and
Kennedy (2001) and Klinger et al. (2001)
reported it from southeast and southwest
France, northern Spain, Austria, the Netherlands,
Zululand (South Africa), Mozambique,
Madagascar, South India, Western
Australia, Chile, Argentina, and California.
It ranges from the upper lower to the upper
upper Maastrichtian (Klinger et al., 2001).
Henderson et al. (1992: 153) noted that this
species was widely distributed but did not
generally range beyond low to midlatitudes.
They remarked that ‘‘it is a useful indicator of
middle to late Maastrichtian age and represents
the last widely distributed heteromorph
taxon to appear in the stratigraphic record.’’ |
| Systematics: |
38
Ordo Ammonoidea
Subordo Ancyloceratina
Superfamilia Turrilitaceae
Familia Baculitidae
Genus Eubaculites
Species Eubaculites carinatus
44
Classis Cephalopoda
Subclassis Nautiloidea
Ordo Ammonoidea
Subordo Ancyloceratina
Superfamilia Turrilitaceae
Familia Baculitidae
Species Eubaculites carinatus
45
Classis Cephalopoda
Subclassis Nautiloidea
Ordo Ammonoidea
Subordo Ancyloceratina
Superfamilia Scaphitaceae
Superfamilia Turrilitaceae
Familia Baculitidae
Familia Scaphitidae
Genus Eubaculites
Species Eubaculites carinatus
46
Ordo Ammonoidea
Subordo Ancyloceratina
Superfamilia Turrilitaceae
Familia Baculitidae
Genus Eubaculites
Species Eubaculites carinatus
48
Ordo Ammonoidea
Subordo Ancyloceratina
Superfamilia Turrilitaceae
Familia Baculitidae
Genus Eubaculites
Species Eubaculites carinatus
|
| Synonym list: |
Cobban & Kennedy (1995):
1986 Eubaculites lyelli d'Orbigny. - Kennedy : p.195 pl. 27, fig. 3-8;
pl. 32, fig. 13, 14 [with full synonymy]
1992 Eubaculites carinatus Morton. - Henderson et al. : p.150 fig. 6F, G, 14-16, 17A-C, G-J, 18, 19 [with additional synonymy]
1993 Eubaculites carinatus Morton. - KLINGER & Kennedy : p.218 fig. 7a-e, 21-30, 31a-g, 32-35, 36e-f, 37-38, 42a, 52g-h
1995 Eubaculites carinatus Morton. - Cobban & Kennedy : p. 21, 22, 24, 25, 2 fig. 14.1, 14.5-14.7, 15.4, 15.6-15.8, 16.13-16.15, 16.23, 16.24, 16.28-16.30, 17.52-17.59, 18.1-18.44
1992 Eubaculites carinatus Morton. - Henderson et al. : p.150 fig. 6f, g, 14-16, 17a-c, g-j, 18, 19 [with additional synonymy]
1993 Eubaculites carinatus Morton. - KLINGER & Kennedy : p.218 fig. 7a-e, 21-30, 31a-g, 32-35, 36e-f, 37-38, 42a, 52g-h
1995 Eubaculites carinatus Morton. - Cobban & Kennedy : p.26 fig. 14.1, 14.5-14.7, 15.4, 15.6-15.18, 16.13-16.15, 16.23, 16.24, 16.28-16.30, 17.52-17.59, 18.1-18.44
1986 Baculites lyelli d'Orbigny. - Kennedy : p.1016 pl. 1, fig. 1-3;
pl. 2, fig. 3-8, 13-21 (with full synonymy)
1986 Baculites lyelli d'Orbigny. - Stinnesbeck : p.207 pl. 9, fig. 6-9;
text-fig. 24D
1987 Baculites lyelli d'Orbigny. - Kennedy : p.195 pl. 27, fig. 5-8;
pl. 32, fig. 13, 14
1992 Eubaculites carinatus Morton. - Henderson et al. : p.150 fig. 6f, g, 14-20 (with additional synonymy)
1987 Eubaculites lyelli d'Orbigny. - Kennedy : p.195 pl. 27, fig. 5-8;
pl. 32, fig. 13, 14 [with full synonymy]
1993 Eubaculites carinatus Morton. - KLINGER & Kennedy : p.218 text-fig. 7a-e, 21-30, 31a-g, 32-35, 36e, f, 37, 38, 42a, 52g, h
2000 Eubaculites carinatus Morton. - Kennedy & Cobban : p.180 pl. 2, fig. 1-23, 27, 28;
text-fig. 3, 4 [with additional synonymy]
1987 Eubaculites lyelli d'Orbigny. - Kennedy : p.195 pl. 27, fig. 5-8;
pl. 32, fig. 13, 14 [with full synonymy]
1993 Eubaculites carinatus Morton. - KLINGER & Kennedy : p.218 text-fig. 7a-e, 21-30, 31a-g, 32-35, 36e, f, 37, 38, 42a, 52g, h
2000 Eubaculites carinatus Morton. - Kennedy & Cobban : p.180 pl. 2, fig. 1-23, 27, 28;
text-fig. 3, 4 [with full synonymy]
|
| Stratigraphy - relative ages: |
upper Maastrichtian - lower Maastrichtian: Ward & Kennedy (1993)
Maastrichtian: Cobban & Kennedy (1995)
Maastrichtian: Kennedy & Cobban (1996)
|
| Specimen: |
Academy of Natural Sciences - Philadelphia, Inventory number: No. 72866
|
| References: |
Morton,S.G.. (1834):
Synopsis of the Organic Remains of the Cretaceous Group of the United States.
d'Orbigny,A. (1847):
Paléontologie, Pls. 1-6 (Geologie Pls. 4-9).
In: M. de Dumont D'Urville, 1846-1 854, Voyage au Pole Sud et dans I'Oceanie sur les corvelles L'Astrolabe et la Zelée pendant les années 1837- 1838-1839-1840 sous le commandément de M. Dumont D'Urville Capitaine du Vaisseau. Pls. 1-9 Eds: Baudry, G..
Conrad,T.A.. (1858):
Observations on a group of Cretaceous fossil shells found in Tippah County, Mississippi, with descriptions of fifty-six new species . Journal of the Academy of Natural Sciences of Philadelphia (2nd ser.) Vol. 3 p. 323-336
Crick,G.C.. (1924):
Appendix A . Transactions of the Geological Society of South Africa Vol. 26 p. 130-140(pl.9)
Stinnesbeck,W.. (1986):
Zu den faunistischen und palökologischen Verhältnissen in der Quriquina Formation (Maastrichtium)Zentral-Chiles . Palaeontographica A194((4-6)) p. 99-237
Kennedy,W.J.. (1986):
Upper Campanian and Maastrichtian ammonites from the Petites-Pyrénées, southern France . Eclogae Geologicae Helvetiae Vol. 79 p. 1001-1037
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The Campanian-Maastrichtian ammonite sequence in the environs of Maastricht (Limburg, the Netherlands) . Newsletters on Stratigraphy Vol. 16 p. 149-168
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The ammonite fauna of the type Maastrichtian with a revision of Ammonites colligatus Binkhorst, 1861. . Bulletin de I 'Institut Royal des Seiences Naturelles de Belgique, Seiences de la Terre Vol. 56 p. 151-267
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The ammonite faunas of the type Maastrichtian, with a revision of Ammonites colligatus Binkhorst, 1861 . Bulletin de l'Institut Royal des Sciences Naturelles de Belgique, Sciences de la Terre Vol. 56 p. 151-267
Henderson,R.A..; Kennedy,W.J.. and McNamara,K.J.. (1992):
Maastrichtian heteromorph ammonites from the Carnarvon Basin, Western Australia . Alcheringa Vol. 16 p. 133-170
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Heteromorph ammonites from the Upper Maastrichtian of Pondicherry, south India . Palaeontology Vol. 35 p. 693-731
KLINGER,H.C.. and Kennedy,W.J.. (1993):
retaceous faunas from Zululand and Natal, South Africa. The heteromorph ammonite genus Eubaculites Spath, 1926. . Annals of the South African Museum, 102:185264
Ward,P.D.. and Kennedy,W.J.. (1993):
Maastrichtian Ammonites from the Biscay Region (France, Spain) . Memoir (The Paleontological Society), Journal of Paleontology34 (Supplement to Vol. 67)(5) p. 1-58
Cobban,W.A.. and Kennedy,W.J.. (1995):
Maastrichtian ammonites chiefly from the Prairie Bluff Chalk in Alabama and Mississippi . Journal of Paleontology Vol. 69(5) p. 1-40
Kennedy,W.J.. and Cobban,W.A.. (1996):
Maastrichtian ammonites from the Hornerstown Formation in New Jersey . Journal of Paleontology Vol. 70(5) p. 798–804
Kennedy,W.J.. and Cobban,W.A.. (2000):
Maastrichtian (Late Cretaceous) ammonites from the Owl Creek Formation on northeastern Missisippi, U.S.A. . Acta Geologica Polonica Vol. 50 p. 175–190
Kennedy,W.J..; GALE,A.S.. and HANSEN,T.A.. (2001):
The last Maastrichtian ammonites from the Brazos River sections in Falls County, Texas . Cretaceous Research Vol. 22 p. 163-171
Landman,N.H..; Johnson,R.O.. and Edwards,L.E.. (2004):
Cephalopods from the Cretaceous/Tertiary Boundary Interval on the Atlantic Coastal Plain, with a Description of the Highest Ammonite Zones in North America, Part 2: Northeastern Monmouth County, New Jersey . Bulletin of the American Museum of Natural History Vol. 287 p. 1-107
Landman,N.H..; Johnson,R.O.. and Edwards,L.E.. (2004):
Cephalopods from the Cretaceous/ Tertiary boundary interval on the Atlantic Coastal Plain, with a description of the highest ammonite zones in North America . Part 1; Maryland. American Museum Novitates Vol. 3454 p. 1-64
Landman,N.H..; Johnson,R.O..; Garb,M.P..; Edwards,L.E.. and Kyte,F.T.. (2007):
Cephalopods from the Cretaceous/Tertiary Boundary Interval on the Atlantic Coastal Plain, with a Description of the Highest Ammonite Zones in North America; Part III; Manasquan River Basin, Monmouth County, New Jersey . Bulletin of the American Museum of Natural History Vol. 303 p. 1-122
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