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Species Catapsydrax africanus Blow & Banner 1962


Diagnosis / Definition:
Pearson et al. (2006):
DESCRIPTION. Type of wall: Cancellate, apparently spinose, ruber/sacculifer-type wall texture. Test morphology: Low trochospiral, slightly lobate test with 3-4 chambers in the final whorl. Chambers inflated, globular, increasing moderately in size with a large inflated bulla covering the umbilicus; in spiral view 3-4 globular chambers increasing moderately in size, sutures straight, moderately depressed; in umbilical view dominated by a large inflated bulla with 3-4 semicircular, infralaminal apertures with a continuous, thickened imperforate rim opening onto each suture which is straight and moderately depressed; in edge view ovoid in shape with bulla showing one aperture opening onto the suture line. Size: Maximum diameter of holotype 0.23 mm, thickness 0.22 mm.
Discussion / Comments:
Pearson et al. (2006):
DISTINGUISHING FEATURES. Catapsydrax africanus is distinguished from C. globiformis by its less spherical test and by having a more inflated bulla that has larger, more open, infralaminal apertures. Catapsydrax howei is larger and has a larger, still more inflated bulla with large circular, infralaminal apertures. DISCUSSION. Blow and Banner (1962) originally placed C. africanus in Globigerinita but this genus is a microperforate taxon, hence it is here placed in Catapsydrax. The species name is changed to agree in gender with the genus Catapsydrax (ICZN, Art. 3 1.2). Catapsydrax africanus, has a low, slightly inflated bulla covering the umbilicus with a single low arched aperture. The infralaminal aperture is bordered by a continuous narrow lip that becomes thickened with gametogenetic calcification. The wall texture of the ruber/sacculifertype also becomes thickened by gametogenetic calcification. The wall texture of C. africanus is the same as in C. unicavus, and the infralaminal apertures are bordered by a continuous, narrow, thickened lip. In C. africanus the bulla is more inflated and the infralaminal apertures are larger. Blow (1979) regarded africanus as a subspecies of Catapsydrax echinatus Bolli, but that species is regarded as an aberrant acarininid in this work (see Berggren and others, Chapter 9, this volume). Catapsydrax africanus is a small form and has apparently been overlooked by many previous workers. We illustrate the holotype in SEM for the first time (Pl. 5.1, Figs. 1-3) as well as newly collected specimens from the type locality (PI. 5.1, Figs. 4-7). PHYLOGENETIC RELATIONSHIPS.- Catapsydrax africanus probably evolved from C. howei in the late middle Eocene by developing a less inflated bulla with smaller infralaminal apertures opening onto the sutures. GEOGRAPHIC DISTRIBUTION.- Known from tropical to mid latitude sites. STABLE ISOTOPE PALEOBIOLOGY. No data available.
Systematics:

22
 Genus Catapsydrax
  Species Catapsydrax africanus

35
  Ordo Foraminiferida
   Superfamilia Globigerinaceae
    Familia Globigerinidae
     Genus Catapsydrax
      Species Catapsydrax africanus
Synonym list:
Spezzaferri & Silva (1990):
1962 Globigerinita africana Blow & Banner. - Blow & Banner : pl. XV; fig. A-C; fig. 11 (i-iv)
1962 Globigerinita africana Blow & Banner. - Blow & Banner :
1990 Catapsydrax africanus Blow & Banner. - Spezzaferri & Silva : p.238 pl. I; fig. 3a
Pearson et al. (2006):
1962 Globigerinita africana Blow & Banner. - Blow & Banner : p.105 pl. 15; fig. A-C (holotype)[Globigerapsis siemi-involuta Zone, Sample FCRM 1645, Lindi, Tanzania.]
1979 Globigerinita echinata africana Blow & Banner. - Blow : pl. 24, fig. 1 and 4; pl. 240, fig. 8 [Zone El4, Lindi, Tanzania]
1979 Globigerinita echinata echinata Bolli. - Blow : pl. 240; fig. 7 [Zone E14, Lindi, Tanzania]. [Not Bolli, 1957b.]
1983 Globigerinita africana Blow & Banner. - Abdel-Kireem : p.88 pl. 2; fig. 14-15 [middle Eocene Truncorotaloides rohri Zone, Mokattam Fm., Egypt]
? 2001 Catapsydrax africana Blow & Banner. - Warraich & Ogasawara : p.43 fig. 12: 1-3 [Zone E10/11, Kirthar Fm., Sulaiman Range, Pakistan].
2006 Catapsydrax africanus Blow & Banner. - Pearson et al. : p.73 pl. 5.1; fig. 1-14 (pl. 5.1; fig. 1-3; new SEMs of holotype of Globigerinita africana Blow and Banner)
Specimen:
Natural History Museum, London, Inventory number: P. 44553
References:

Blow,W.H. and Banner,F.T. (1962):
The Mid-Tertiary (Upper Eocene to Aquitanian) Globigerinaceae.
In: Fundamentals of Mid‑Tertiary Stratigraphical Correlation Eds: Eames, F.E.Banner, F.T.Blow, W.H.Clarke, W.J. p. 61‑151

Blow,W.H. (1979):
The Cainozoic Globigerinida. 3 Vols p. 1413 pp

Abdel-Kireem,M.R.. (1983):
Planktonic foraminifera of Mokattam Formation (Eocene) of Gebel Mokattam, Cairo, Egypt . Revue de Micropaleontologie Vol. 28 p. 77-96

Spezzaferri,S. and Silva,I.S.. (1990):
Oligocene planktonic foraminiferal biostratigraphy and paleoclimatic interpretation from Hole 538A, DSDP Leg 77, Gulf of Mexico . Palaeogeography, Palaeoclimatology, Palaeoecology Vol. 83 p. 217-263

Warraich,M.Y.. and Ogasawara,K.. (2001):
Tethyan Paleocene-Eocene planktic foraminifera from the Rakhi Nala and Zinda Pir land sections of the Sulaiman Range, Pakistan . Science Reports of the Institute of Geoscience University of Tsukuba, section B Vol. 22 p. 1-59

Pearson,P.N.; Olsson,R.K.; Hemleben,C.; Huber,B.T. and Berggren,W.A. (2006):
Atlas of Eocene Planktonic Foraminifera. p. 1-513
Anonymous: Unedited TaxonConcept data
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