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Species Subbotina gortanii Borsetti 1959


Diagnosis / Definition:
Pearson et al. (2006):
DESCRIPTION. Type of wall: Cancellate, normal perforate, spinose, ruber/sacculifer-type wall texture. Test morphology: High trochospiral, globular in outline, chambers globular; in spiral view 4 globular, loosely embracing chambers in ultimate whorl, increasing moderately in size, sutures deeply depressed, straight; in umbilical view 4 globular, loosely embracing chambers, increasing moderately in size, sutures deeply depressed, straight, umbilicus large, enclosed by surrounding chambers, aperture umbilical, bordered by a thickened, narrow rim; in edge view chambers arranged in a high, loosely coiled spire. Size: Maximum diameter of holotype 0.60 mm, thickness 0.63 mm.
Discussion / Comments:
Spezzaferri & Silva (1990):
Very rare.
Pearson et al. (2006):
DISTINGUISHING FEATURES.- The species is characterized by its trochospiral loosely coiled test, globular chambers, and umbilical aperture. DISCUSSION.- Blow (1979) regarded Globigerina gortanii gortanii as the senior synonym of Globigerina turritilina turritilina Blow and Banner, but retained the separation of this form from Globigerina turritilina praeturritilina Blow and Banner. Blow and Banner (1 962) separated Globigerina turritilina turritilina from Globigerina turritilina praeturritilina on the basis of slightly less inflated chambers, a broader umbilicus, and a higher aperture in G. turritilina praeturritilina. The holotypes of Blow and Banner's subspecies are illustrated for the first time in SEM on Plate 6.10, Figs. 9-12 and show little difference between the two morphotypes in these characters. Khalilov's Globigerina pseudocorpulenta is a high-spired form that Blow (1979) considered a possible senior synonym of praeturritilina. The holotype was viewed by WAB who was unable to place the specimen unequivocally, hence we hold it in questionable synonymy with gortanii but recommend suppression of the name. PHYLOGENETIC RELATIONSHIPS.- Subbotina gortanii appears related to Subbotina yeguaensis, which has a moderately elevated spire of loosely embracing, globular chambers and a ruber/sacculifer-type wall texture. An increase in the height of the coil would lead to the development of the gortanii morphology. GEOGRAPHIC D1STNBUTION.- Apparently global in low to mid latitudes. STABLE ISOTOPE PALEOBIOLOGY.- Pearson and others (2001) recorded this species (as S. cf. S. praeturritilina) as a deep dwelling form in the upper Eocene. However Douglas and Savin (1978) and Boersma and others (1987) recorded relatively negative ň18O for this species from the Oligocene, suggesting that it might have migrated into shallower water habitats.
Systematics:

22
 Species Subbotina gortanii
  Species Turborotalia increbescens
   Species Turborotalia pseudoampliapertura

35
  Ordo Foraminiferida
   Superfamilia Globigerinaceae
    Familia Globigerinidae
     Genus Subbotina
      Species Subbotina gortanii
Synonym list:
Spezzaferri & Silva (1990):
1959 Catapsydrax gortanii Borsetti. - Borsetti :
1959 Catapsydrax gortanii Borsetti. - Borsetti : pl. 1; fig. 1a-1c
1990 Subbotina gortanii Borsetti. - Spezzaferri & Silva : p.258 pl. XVI; fig. 3a-d
Pearson et al. (2006):
? 1956 Globigerina pseudocorpulenta Khalilov. - Khalilov : p.246 pl. 4; fig. 3a-c [upper Eocene, Maly Caucasus]
1959 Catapsydrax gortanii Borsetti. - Borsetti : p.205 pl. 1; fig. 1a-d (1a-c = holotype) [lower Oligocene, Piacenza Province, northern Italy]
1962 Globigerina turritiliana turritiliana Blow & Banner. - Blow & Banner : p.98 pl. 13; fig. D-G [lower Oligocene Globigerina oligocaenica Zone, Lindi area, Tanzania]
1962 Globigerina turritiliana praeturritiliana Blow & Banner. - Blow & Banner : p.99 pl. 13; fig. A-C [upper Eocene Globigerapsis semiinvoluta Zone, Lindi area, Tanzania]
1969 Globigerina gortanii gortanii Borsetti. - Blow : p.320 pl. 17; fig. 1 [lower Oligocene Zone 02, Sample FCRM 1964, Lindi area, Tanzania]
1979 Globigerina gortanii gortanii Borsetti. - Blow : p.851 pl. 17, fig. 1 [reillustration of Blow, 1969, pl. 17: fig. l]; pl. 247, fig. 4 [lower Oligocene Zone 0213, DSDP Site 14, central South Atlantic Ocean]; pl. 251, fig. 9 [upper Oligocene Zone 06, DSDP Site 14, central South Atlantic Ocean]
1980 Globigerina gortanii gortanii Borsetti. - Charollais et al. : p.62 pl. 5; fig. 1, 2 [lower Oligocene, France]
1983 Globigerina gortanii Borsetti. - Pujol : p.651 pl. 11, fig. 4 [lower Oligocene Zone 03, DSDP Hole 5 16F, Rio Grande Rise, southwest South Atlantic Ocean]; pl. 15, fig. 4 [Zone N516, DSDP Site 516, Rio Grande Rise, southwest South Atlantic Ocean]
1985 Globigerina gortanii Borsetti. - Snyder & Waters : p.459 pl. 1; fig. 16-18 [upper Oligocene Zone 05, DSDP Hole 549A, Goban Spur, eastern North Atlantic Ocean]
2006 Subbotina gortanii Borsetti. - Pearson et al. : p.136 pl. 10; fig. 1-17 (Pl. 6.1 0, Figs. 1-3: new SEMs of holotype of Globigerina turritilina turritilina Blow and Banner); (Pl. 6.10, Figs. 9- 12: new SEMs of holotype of Globigerina turritilina praeturritilina Blow and Banner)
Specimen:
Institute of Geology and Paleontology, University of Bologna, Italy, Inventory number: IF - 376
References:

Khalilov,D.M. (1956):
O pelagicheskoy faune foraminifer Paleogenovykh otlozheniy Azerbaydzhana . Trudy Instituta Geologii, Akademiya Nauk Azerbaydzhanskoy SSR Vol. 17

Borsetti,A.M. (1959):
Tre nouvi Foraminiferi planctonici dell'Oligocene piacentino . G. Geol. Vol. 27 p. 205-212

Blow,W.H. and Banner,F.T. (1962):
The Mid-Tertiary (Upper Eocene to Aquitanian) Globigerinaceae.
In: Fundamentals of Mid‑Tertiary Stratigraphical Correlation Eds: Eames, F.E.Banner, F.T.Blow, W.H.Clarke, W.J. p. 61‑151

Blow,W.H. (1969):
Late middle Eocene to Recent planktonic foraminiferal biostratigraphy.
In: Proceedings of the First International Conference on Planktonic Microfossils, Geneva 1967 Vol. 1 Eds: Bronnimann, P.Renz, H.H. p. 199-422

Blow,W.H. (1979):
The Cainozoic Globigerinida. 3 Vols p. 1413 pp

Charollais,J..; Hochuli,P..; Oektlt,H..; Perch-Nielsen,K.; Toumarkine,M.; Rögl,F. and Pajris,-.J.. (1980):
Les Marnes Foraminiféres et les Schistes a Meletta des Chaines Subalpines septentrionales (Haute-Savoie, France) . Eclogae Geologicae Helvetiae Vol. 73 p. 9-69

Pujol,C. (1983):
Cenozoic Planktonic Foraminiferal Biostratigraphy of the Southwestern Atlantic (Rio Grande Rise): Deep Sea Drilling Project Leg 72.
In: Initial Reports of the Deep Sea Drilling Project Vol. 72 Eds: Barker, P.F.Johnson, D.A. p. 623-673

Snyder,S.W. and Waters,V.J. (1985):
Cenozoic planktonic foraminiferal biostratigraphy of the Goban Spur region.
In: Deep Sea Drilling Project Vol. 80

Spezzaferri,S. and Silva,I.S.. (1990):
Oligocene planktonic foraminiferal biostratigraphy and paleoclimatic interpretation from Hole 538A, DSDP Leg 77, Gulf of Mexico . Palaeogeography, Palaeoclimatology, Palaeoecology Vol. 83 p. 217-263

Pearson,P.N.; Olsson,R.K.; Hemleben,C.; Huber,B.T. and Berggren,W.A. (2006):
Atlas of Eocene Planktonic Foraminifera. p. 1-513
Anonymous: Unedited TaxonConcept data
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