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Species Subbotina hagni Gohrbandt 1967



Diagnosis / Definition:
Pearson et al. (2006):
DESCRIPTION. Type of wall: Cancellate, normal perforate, spinose, ruber/sacculifer-type wall texture. Test morphology: Test low trochospiral, quadrate in outline, chambers globular; in spiral view 4 globular, somewhat embracing chambers in ultimate whorl, increasing moderately in size, sutures moderately depressed, straight; in umbilical view 4-4 1/2 globular, somewhat embracing chambers, increasing moderately in size, sutures moderately depressed, straight, umbilicus small, enclosed by surrounding chambers, aperture umbilical to extraumbilical, bordered by a thin, irregular lip; in edge view chambers globular in shape, embracing; aperture visible as a low arch, bordered by a thin, irregular lip. Size: Maximum diameter of holotype 0.58 mm, thickness 0.34 mm.
Discussion / Comments:
Pearson et al. (2006):
DlSTINGUISHING FEATURES.- The species is characterized by its generally large adult size, quadrate test, and globular, embracing, chambers with a low- arched, umbilical-extraumbilical aperture bordered by a thin irregular lip. DISCUSSTON.- Gohrbandt (1 967) noted that the illustrations of Globigerina eocaena Guembel by Subbotina (1953) corresponded to his new species, although he did not formally place these in synonomy with his new species. He also acknowledged that Hagn and Lindenberg (1966), who selected a neotype for G. eocaena, did not consider Subbotina's illustrations of G. eocaena to be correctly identified. Subbotina hagni has not been widely recorded by workers, perhaps because it has been generally grouped with S. eocuena populations. We have identified it in the Aragon Formation of Mexico and California, which indicates a widespread distribution for this species. PHYLOGENETIC RELATIONSHIPS.- Subbotina hagni belongs to a group of closely related Eocene large-sized subbotinids, which includes S. eocaena and S. corpulenta. It probably evolved from S. eocaena by an increase in number of chambers in the ultimate whorl and the development of a more quadrate-shaped test. GEOGRAPHIC DISTRIBUTION.- Apparently global in low to mid latitudes. STABLE ISOTOPE PALEOBIOLOGY.- No data available
Systematics:

35
 Ordo Foraminiferida
  Superfamilia Globigerinaceae
   Familia Globigerinidae
    Genus Subbotina
     Species Subbotina hagni
Synonym list:
Pearson et al. (2006):
1953 Globigerina eocaena Guembel. - Subbotina : p.85 pl. 6, fig. 5a-c; pl. 7, fig. 1a-c [upper Eocene Acarinina Zone (upper part), Kuban River, northern Caucasus]. [Not Guembel, 1868.]
1967 Globigerina hagni Gohrbandt. - Gohrbrandt : p.324 pl. 1; fig. 1-9 (1-3 = holotype, 4-9 = paratypes), [middle Eocene, Helvetikum, Salzburg, Austria]
1977 Subbotina hagni Gohrbandt. - Poore & Brabb : p.269 pl. 4; fig. 13, 14 [middle Eocene Zone E 1 211 3, Butano Sandstone, Santa Cruz Mountains, California]
1985 Globigerina hagni Gohrbandt. - Toumarkine & Luterbacher : p.150 fig. 42-7a-c (reiilustration of holotype); fig. 8, 9 [middle Eocene Hantkenina nuttalli Zone, Possagno, Italy]
2001 Subbotina hagni Gohrbandt. - Warraich & Ogasawara : p.48 fig. 14: 5-7 [middle Eocene, Baska Fm., Sulaiman Range, Pakistan].
2006 Subbotina hagni Gohrbandt. - Pearson et al. : p.139 pl. 6.11; fig. 1-17 (Pl. 6.11, Figs. 1-3: new SEMs of topotype of Globigerina hagni Gohrbandt)
Specimen:
Naturhistorisches Museum, Vienna, Austria, Inventory number: 683/10/1966
References:

Subbotina,N.N. (1953):
Iskopaemye foraminifery SSSR (Globigerinidy, Khantkenininidy i Globorotaliidy) . Trudy Vsesoyznogo Nauchno-Issledovatel'skogo Geologo-razvedochnogo Instituta (VNIGRI) Vol. 76 p. 296

Gohrbrandt,K.H.A. (1967):
Some new foraminiferal species from the Austrian Eocene . Micropaleontolgy Vol. 13 p. 319-326

Poore,R.Z. and Brabb,E.E.. (1977):
Eocene and Oligocene planktonic foraminifera from the Upper Butano Sandstone and type San Lorenzo Formation, Santa Cruz Mountains, California . Journal of Foraminiferal Research Vol. 7 p. 249-272

Toumarkine,M. and Luterbacher,H.P. (1985):
Paleocene and Eocene Planktic Foraminifera.
In: Plankton Stratigraphy p. 87-154

Warraich,M.Y.. and Ogasawara,K.. (2001):
Tethyan Paleocene-Eocene planktic foraminifera from the Rakhi Nala and Zinda Pir land sections of the Sulaiman Range, Pakistan . Science Reports of the Institute of Geoscience University of Tsukuba, section B Vol. 22 p. 1-59

Pearson,P.N.; Olsson,R.K.; Hemleben,C.; Huber,B.T. and Berggren,W.A. (2006):
Atlas of Eocene Planktonic Foraminifera. p. 1-513

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