Turborotalita carcoselleensis Toumarkine & Bolli 1975 (TaxonConcept

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Species Turborotalita carcoselleensis Toumarkine & Bolli 1975

Diagnosis / Definition:

Pearson et al. (2006):
DESCRIPTION. Type of wall: Normal perforate, spinose, rubertype, cancellate wall structure, covered by a crust in adult stage. Test morphology: Test low trochospiral, slightly lobulate in outline, chambers globular; in spiral view 4 1/2 -5 globular, slightly embracing chambers in ultimate whorl, increasing moderately in size, ultimate chamber often reduced in size, sutures moderately depressed, straight; in umbilical view 4 1/2-5 globular, slightly embracing chambers, increasing moderately in size, ultimate chamber often reduced and directed towards and over the umbilicus, sutures moderately depressed, straight, umbilicus small and often covered by ultimate chamber, aperture umbilical, a wide arch bordered by an imperforate rim or narrow thickened lip; in edge view chambers globular in shape, slightly embracing. Size: Maximum diameter of holotype 0.27 mm, maximum width of thickness 0.19 mm.

Discussion / Comments:

Pearson et al. (2006):
DISTINGUISHING FEATURES.- Turborotalita carcoselleensis is characterized by its small, slightly lobulate test, the umbilically directed ultimate chamber, and the ruber-type cancellate wall texture. DISCUSSION.- There has been considerable confusion regarding the identification of Globigerina wilsoni Cole. Toumarkine and Luterbacher (1985) reillustrated the line drawing of the holotype of Globigerinn wilsoni along with three SEM images they identified as that species from the type Guayabal Formation, from where G. wilsoni was described. New SEM images ofthe holotype of G. wilsoni (Pl. 6.19, Figs. 15- 17), although poorly preserved, show that it differs from the hypotypes illustrated by Toumarkine and Luterbacher in that the ultimate chamber is more flattened and extends towards and over the umbilicus. In this respect, the holotype appears much more like Turborotalia carcoselleensis (Toumarkine and Bolli) (=Turborotalita carcoselleensis), which is common in the Guayabal Formation. However, the poor state of preservation does not allow an accurate identification, and we place wilsoni in questionable prior synonymy with carcoselleensis. The hypotypes ofwilsoni illustrated by Toumarkine and Luterbacher are placed in Parasubbotina pseudowilsoni n. sp. (see discussion under this species in chapter 5). Turborotalita carcoselleensis differs from P pseudowilsoni n. sp., which is described from the Guayabal Formation, in the more moderate increase in chamber size and the distinctive ultimate chamber that extends as a narrow, reduced chamber over the umbilicus. PHYLOGENETIC RELATIONSHIPS.- The origin of Turborotalita carcoselleensis is uncertain. Morphologically it appears closest to Globoturborotalita in the arrangement of chambers around a small umbilicus and in the umbilical aperture. In addition, species of Globoturborotalita are small in size, as is Turborotalita. Globoturborotalita bassriverensis n. sp. is a possible ancestor. GEOGRAPHIC DISTRIBUTION .- Widespread in low to middle latitudes. STABLE ISOTOPE PALEOBIOLOGY.- No data available.

Systematics:

35
Ordo Foraminiferida
  Superfamilia Globigerinaceae
   Familia Globigerinidae
    Genus Turborotalita
     Species Turborotalita carcoselleensis

Synonym list:

Pearson et al. (2006):

? 1927 Globigerina wilsoni Cole. - Cole : p.34 pl. 4; fig. 8-9 [middle Eocene Zone E10/11, Guayabal Fm., Tampico, Mexico]

1975 Globorotaloides carcoselleensis Toumarkine & Bolli. - Toumarkine & Bolli : p.81 pl. 5; fig. 23-28 (fig. 24 = holotype); [middle Eocene Globorotalia cerroazulensis possagnoensis Zone, Possagno, northern Italy]

1975 Globorotaloides carcoselleensis Toumarkine & Bolli. - Toumarkine : p.742 pl. 1; fig. 20-21 [middle Eocene Morozovella lehneri Zone, DSDP Site 3 13, central Pacific Ocean]

1977 Globorotaloides carcoselleensis Toumarkine & Bolli. - Poore & Brabb : p.260 pl. 5, fig. 4, 5 [middle Eocene Zone E13, Twobar Shale Member, San Lorenzo Fm., Santa Cruz Mountains, California]; pl. 7, fig. 1, 2 [upper Eocene Zone E14/16, Twobar Shale Member, San Lorenzo Fm., Santa Cruz Mountains, California]

1978 Globorotaloides carcoselleensis Toumarkine & Bolli. - Toumarkine : p.712 pl. 7; fig. 13-18 [middle Eocene Orbulinoides beckmanni Zone, DSDP Site 363, Walvis Ridge, South Atlantic Ocean]

p 1985 Turborotalia wilsoni Cole. - Toumarkine & Luterbacher : p.126 fig. 27.1 (partim; not fig. 27.2-4); (reillustration of holotype).

1985 Globorotaloides carcoselleensis Toumarkine & Bolli. - Toumarkine & Luterbacher : p.150 fig. 41.9-16 (fig. 41.9, 10, reillustration of holotype and paratype), fig. 41.11, 12a-b, 15 [middle Eocene Orbulinoides beckmanni Zone, DSDP Site 363, Walvis Ridge, South Atlantic Ocean (figs. 13, 16 from Toumarkine, 1975, pl. l)]; fig. 41.13, 14, 16 [middle Eocene Morozovella lehneri Zme, DSDP Site 3 13, central Pacific Ocean (figs. ll, 12a-b, 15 from Toumarkine, 1978, pl. 7)]

1988 Globorotaloides carcoselleensis Toumarkine & Bolli. - Poore & Bybell : p.18 pl. 1; fig. 4, 5, 7 [middle Eocene Morozovella lekneri Zone, Shark River Fm., ACGS #4 Borehole, New Jersey Coastal Plain]

2006 Turborotalita carcoselleensis Toumarkine & Bolli. - Pearson et al. : p.155 pl. 6.19; fig. 1-17 (Pl. 6.19, Figs. 1-3: reillustration of holotype and paratype of Globorotaloides carcoselleensis Toumarkine and Bolli); (PI. 6.19, Figs. 15-17: new SEMS of holotype of Globigerina wilsoni Cole)

Specimen:

National Museum of Natural History, Basel, Inventory number: C27330

References:

Cole,W.S. (1927):
A foraminiferal fauna from the Guabal Formation in Mexico . Bulletins of American paleontology Vol. 14 p. 1-46

Toumarkine,M. (1975):
Middle and Late Eocene planktonic foraminifera from the northwestern Pacific, Leg 32 of the Deep Sea Drilling Project.
In: Initial Reports of the Deep Sea Drilling Project Vol. 32 Eds: Larson, R.L.Moberly, R. p. 735-751

Toumarkine,M. and Bolli,H.M. (1975):
Foraminifères planctoniques de l'Eocène moyen et supérieur de la coupe de Possagno . Schweizerische Paläontologische Abhandlungen Vol. 97 p. 69-185

Poore,R.Z. and Brabb,E.E.. (1977):
Eocene and Oligocene planktonic foraminifera from the Upper Butano Sandstone and type San Lorenzo Formation, Santa Cruz Mountains, California . Journal of Foraminiferal Research Vol. 7 p. 249-272

Toumarkine,M. (1978):
Planktonic Foraminiferal Biostratigraphy of the Paleogene of Sites 360 to 364 and the Neogene of Sites 362A, 363, and 364 Leg 40.
In: Initial Results of the Deep Sea Drilling Project Vol. 40

Toumarkine,M. and Luterbacher,H.P. (1985):
Paleocene and Eocene Planktic Foraminifera.
In: Plankton Stratigraphy p. 87-154

Poore,R.Z. and Bybell,L.. (1988):
Eocene to Miocene biostratigraphy of New Jersey Core ACGS # 4: Implications for regional stratigraphy . U. S. Geological Survey Bulletin 1829 p. 1-22

Pearson,P.N.; Olsson,R.K.; Hemleben,C.; Huber,B.T. and Berggren,W.A. (2006):
Atlas of Eocene Planktonic Foraminifera. p. 1-513

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