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Species Brahmaites (Anabrahmaites) vishnu Forbes 1846

[1]
[2]
[1] text-fig. 8a-b Ifrim et al. (2004) [2] text-fig. 9a-d Ifrim et al. (2004)

Alternative name:
Brahmaites vishnu Forbes 1846
Diagnosis / Definition:
Ifrim et al. (2004):
Description: Moderately evolute with intermediate expansion rate. The broadly rounded venter grades into rounded flanks with maximum breadth above the middle flanks. Flanks bend narrowly into a steep overhanging short umbilical wall. Ornament on internal moulds is restricted to three constrictions per whorl, which are associated with two collar ribs. Ribs and constrictions are parallel. They are straight and prorsiradiate on the flanks, and cross the venter in a wide arch. On two examples with goethitized conchs ribs superimpose the constrictions. The suture line is moderately complex, E is approximately as deep as the trifid L. Bifid LIE and UIL become subdivided in larger specimens. The sutural lobe is retracted. Its length increases strongly with the growing number of umbilical lobes and the grade of incision.
Ward & Kennedy (1993):
Description. - Holotype of Puzosia haugi is fragmentary composite mold with original estimated diameter of 63-65 mm. Coiling very evolute, with U = estimated 47 percent of diameter and around 30 percent of previous whorl being covered. Umbilicus of moderate depth, with flattened subvertical wall. Whorl section depressed, with whorl breadth to height ratio of 1.16. Umbilical shoulder broadly rounded, flanks rounded, outer flanks converging to broadly rounded venter. Estimated four narrow, deep constrictions per whorl on inner whorls to diameter of 30 mm. They are straight and prorsiradiate, and flanked by narrow collar-ribs that may be incipiently bullate; flanks otherwise smooth. Two constrictions on outer whorl separated by arc of 100° approximately, straight and prorsiradiate on umbilical wall, cutting deep into umbilical shoulder and prorsiradiate on flank, flexing slightly forwards and concave on outer flank and crossing venter in broad convexity with linguoid peak over siphonal region. Adapical collar rib stronger than adapertural one and bears pronounced bulla in association with last-preserved constriction, while adapical rib associated with first constriction bears siphonal tubercle. (Region damaged at end of specimen.) Feeble and irregular primary ribs present on first part of outer whorl. Nine primary ribs on last part narrow and rounded, and separated by somewhat wider interspaces. Ribs arise on upper part of umbilical shoulder, strengthened and prorsiradiate on innermost flank but flexed back, straight and only feebly prorsiradiate on mid- to outer flank. They weaken markedly on ventrolateral shoulder and efface over siphonal region.
Discussion / Comments:
Ifrim et al. (2004):
Remarks: The umbilicus increases during growth from 27 per cent at a diameter of 8·4 mm to 35 per cent at a diameter of 16·5 mm. The absence ofumbilical bullae and a depressed whorl section differentiate B. (A.) vishnu from Brahmaites (B.) brahma (Forbes, 1846). Occurrence: Brahmaites (Anabrahmaites) vishnu occurs in the Maastrichtian of southern India (Kennedy and Henderson 1992b), Armenia (Atabekian and Akopian 1970), Madagascar (Collignon 1938), south-west France and northern Spain (Ward and Kennedy 1993). In the US Gulf Coast area the genus Brahmaites was first described by Cobban and Kennedy (1991a) from the Nacatotch Sand, Arkansas, but these specimen have not been determined to species level.
Ward & Kennedy (1993):
Types. - Syntypes are BMNH C5 1026-5 1028, including the original of Forbes, 1846, Pl. 7, fig. 9 (C51026), from the Valudavur Formation of Pondicherry, south India. The holotype, by monotypy, of Puzosia haugi Seunes, 189 1 (p. 20, Pl. 15(6), fig. 1) is an unregistered specimen in the Sorbonne Collections (now in the UniversitC Paris VI), from the "Assize a Pachydiscus Fresvillensis, P. Jacquoti et a Stegaster," Route de Gan a Rébénacq, Pyrénées-Atlantiques, France (Figure 25.1-25.3). Discussion. - We can see no significant differences between the holotype of Puzosia haugi (a composite mold) and specimens of Brahmaites (A.) vishnu from south India, examples of which are illustrated for comparison in Figure 26.1, 26.6-26.8. The extent of the early smooth stage is somewhat variable, being rather longer in the French specimen than some Indian examples, but we take this to be no more than intraspecific variation. The specimen from Tercis referred to Gaudryceras planorbiforme of Bohm by de Grossouvre (1894, Pl. 27, fig. 2) is a gaudryceratid. The specimen from near Alcoy, Spain (de Grossouvre, 1894, Pl. 35, fig. 7) is a B. (A.) vishnu a little larger than the present specimen with coarse ribs, as in some Indian specimens of the same size. We are uncertain of the affinities of Brahmaites mikobokensis Collignon, 197 1 (p. 22, Pl. 647, figs. 2400, 2401), which lacks siphonal tubercles in internal molds from Zululand. Occurrence. - Upper Maastrichtian of Pyrénées-Atlantiques, France. Imprecisely dated near Alcoy, Spain; Maastrichtian of Armenia and Madagascar.
Systematics:

36
 Ordo Ammonoidea
  Subordo Ammonitina
   Superfamilia Desmocerataceae
    Familia Kossmaticeratidae
     Subfamilia Kossmaticeratinae
      Genus Brahmaites
       Subgenus Brahmaites (Anabrahmaites)
        Species Brahmaites (Anabrahmaites) vishnu

38
  Ordo Ammonoidea
   Subordo Ammonitina
    Superfamilia Desmocerataceae
     Familia Kossmaticeratidae
      Subfamilia Kossmaticeratinae
       Genus Brahmaites
        Species Brahmaites (Anabrahmaites) vishnu
Synonym list:
Ifrim et al. (2004):
1846 Ammonites vishnu Forbes. - Forbes : p.100 pl. 7, fig. 9
1938 Brahmaites haugi Seunes. - COLLIGNON : p.45 pl. 7, fig. 3
1970 Brahmaites vishnu Forbes. - ATABEKIAN & AKOPIAN : p.37 pl. 2, fig. 6
1992 Brahmaites (Anabrahmaites) vishnu Forbes. - Kennedy & Henderson : p.418 pl. 6, figs. 25-26; pl. 9, figs. 5-7, 17-20; pl. 10, fig. 5; pl. 17, figs. 8, 10-11 (with full synonymy)
1993 Brahmaites (Anabrahmaites) vishnu Forbes. - Kennedy & Hancock : p.580 pl. 1, figs. 5-6
2004 Brahmaites (Anabrahmaites) vishnu Forbes. - Ifrim et al. : 20, 21 text-figs. 8a-b, 9a-d
Ward & Kennedy (1993):
1846 Ammonites vishnu Forbes. - Forbes : p.100 pl. 7; fig. 9
1865 Ammonites vishnu Forbes. - Stoliczka : p.164 pl. 79; fig. 5
1891 Puzosia haugi Seunes. - Seunes : p.20 pl. 15(6); fig. 1
1894 Gaudryceras sp. planorbiforme Böhm. - Grossouvre : p.231 non pl. 27, fig. 2; pl. 34, fig. 4, 5; pl. 35, fig. 7
1897 Brahmaites sp. vishnu Forbes. - Kossmat : 46(153) pl. 8(19); fig. 10
1925 Brahmaites vishnu Forbes. - Diener : p.103
1938 Brahmaites haugi Seunes. - COLLIGNON : p.100 pl. 7; fig. 3
1970 Brahmaites vishnu Forbes. - ATABEKIAN & AKOPIAN : p.37 pl. 2; fig. 1
1993 Brahmaites (Anabrahmaites) vishnu Forbes. - Ward & Kennedy : p. 27, 28 fig. 25.1-25.3, 26.1, 26.6-26.8
Stratigraphy - relative ages:
Maastrichtian: Ifrim et al. (2004)
upper Maastrichtian - upper Maastrichtian: Ward & Kennedy (1993)
References:

Forbes,E.. (1846):
Report on the Cretaceous fossil invertebrates from southern India, collected by Mr. Kaye and Mr. Cunliffe . Transactions of the Geological Society of London Vol. 2(7) p. 97–174

Stoliczka,F.. (1865):
The fossil Cephalopoda of the Cretaceous rocks of southern India. Ammonitidae with revision of the Nautilidae etc. . Memoirs of the Geological Survey of India, (1), Palaeontologica Indica6-9 p. 107-154

Seunes,J.. (1891):
Contributions à l’étude des Céphalopodes du Crétacé Supérieur de France. Ammonites du Calcaire à Baculites du Cotenin (Suite); II. Ammonites du Campanien de la région sous-Pyrénéenne, Département de Landes . Mémoirs de la Société Géologique de France, Paléontologie Vol. 2(2) p. 8-22

Grossouvre,d.A.. (1894):
Recherches sur la Craie Supérieure, 2, Paléontologie. Les ammonites de la Craie Supérieure, Mémoirs pour Servir à l’Explication de la Carte géologique détaillée de la France (mis-dated 1893).. p. 264

Kossmat,F.. (1897):
Untersuchungen über die Südindische Kreideformation. Zweiter Theil . Beiträge zur Paläontologie Österreich-Ungarns und des Orient Vol. 11 p. 1-46 (108-153)

Diener,C.. (1925):
Ammonoidea neocretacea . Fossilium Cat, (1: Animalia) Vol. 29 p. 244 pp.

COLLIGNON,M.. (1938):
Ammonites Campaniennes et Maastrichtiennes de I'ouest et du sud de Madagascar. . Annales Geologiques du Service des Mines de Madagascar Vol. 9 p. 55-118

ATABEKIAN,A.A.. and AKOPIAN,v.T.. (1970):
Late Cretaceous ammonites of the Armenian SSR (Pachydiscidae, Kossmaticeratidae and Scaphitidae) . Proceedings of the Armenian Academy of Science, Earth Seiences [In Georgian] Vol. 5 p. 31-42

Kennedy,W.J.. and Henderson,R.A.. (1992):
Non-heteromorph ammonites from the Upper Maastrichtian of Pondicherry, south India . Palaeontology Vol. 35 p. 381-442

Kennedy,W.J.. and Hancock,J.M.. (1993):
Upper Maastrichtian ammonites from the Marnes de Nay between Gan and Rebenacq (Pyrenees-Atlantiques), France . Geobios Vol. 26 p. 575-594

Ward,P.D.. and Kennedy,W.J.. (1993):
Maastrichtian Ammonites from the Biscay Region (France, Spain) . Memoir (The Paleontological Society), Journal of Paleontology34 (Supplement to Vol. 67)(5) p. 1-58

Ifrim,C..; Stinnesbeck,W.. and López-Oliva,J.G.. (2004):
Maastrichtian cephalopods from Cerralvo, north-eastern Mexico . Palaeontology Vol. 47(6) p. 1575–1627
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