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Species Morozovelloides bandyi Fleisher 1974


Diagnosis / Definition:
Pearson et al. (2006):
DESCRIPTION. Type of wall: Muricate, normal perforate with smooth areas on the spiral side. Test morphology: Planoconvex to lenticular test, coiled in a low trochospiral, with concentration of bladed muricae around the petaloid periphery that constitute a discontinuous muricocarina. Four to five chambers in the final whorl, added in such a way that their long axes are strongly disjunct when viewed on the dorsal side. Typically the chambers are imbricate, sometimes strongly so, as if tilted relative to the dorso-ventral plane. Small supplementary apertures are commonly present at the base of the sutures between the penultimate and antepenultimate chambers, and more rarely earlier in the spire. Primary aperture is a low, flat arch, in an umbilical-extraumbilical position. Umbilicus moderately broad and deep. Ventral sutures depressed with umbilical shoulders of chambers commonly showing concentrations of muricae (but not to the extent seen in M. coronatus, see below). Size: Maximum diameter of holotype about 0.28 mm; height 0.16 mm.
Discussion / Comments:
Pearson et al. (2006):
DISTINGUISHING FEATURES.- Distinguished from all other species of Morozovelloides by the strongly disjunct long axes of the chambers, as seen in spiral view, and from Acarinina praetopilensis by having a more dorso-ventrally flattened test and peripheral (as opposed to circum-cameral) muricocarina. DISCUSSION.- M orozovelloides bandyi is a common form in Zones E8 and E9, but first appears in the upper part of Zone E7 where it intergrades with Acarinina praetopilensis. It has probably been described previously on many occasions as crassata or spinulosa, but such synonymies in the older literature are difficult to verify because M. bandyi is strongly homeomorphic to the Morozovella aequa-M. subbotinae group of the lower Eocene. PHYLOGENETIC RELATIONSHIPS.- Morozovelloides bandyi evolved from Acarinina praetopilensis and gave rise to M. crassatus. STRATIGRAPHIC RANGE.- First appears in the upper Part of Zone E72 intergrading with A. praetopilensis. Most common in Zones E8 and E9, but ranging as high as Zone E10. GEOGRAPHIC DISTRIBUTION.- Cosmopolitan. STABLE ISOTOPE PALEOBIOL0GY.- Oxygen and carbon isotopes indicate a mixed layer, phososymbiotic habitat (Boersma and others, 1987 [recorded as Morozovella cf. aequa], Pearson and others, 1993 and Pearson and others, 2001 [recorded as Morozovella crassata]). Boron isotope data suggest a deeper habitat than co-occurring M. crassatus (Pearson and Palmer, 1999, recorded as Morozovella crassata; Morozovelloides crassatus recorded as Morozovella spinulosa). REPOSITORY.- Deposited in the Cushman Collection at the Smithsonian Museum of Natural History, Washington, D.C.
Systematics:

35
 Ordo Foraminiferida
  Superfamilia Globigerinaceae
   Familia Truncorotaloididae
    Genus Morozovelloides
     Species Morozovelloides bandyi
Synonym list:
Pearson et al. (2006):
1930 Globorotalia crassata Cushman. - Cushman & Barksdale : p. 67-68 pl. 12; fig. 7a-b [Eocene, Martinez Fm., California]
1939 Globorotalia crassata Cushman. - Cushman : p.74 pl. 12; fig. 19 ["upper Eocene", submarine core off eastern north America]; [Not Cushman, 1925]
1953 Globorotalia spinulosa Cushman. - Beckmann : p. 397-398 pl. 26; fig. 13 [middle upper Eocene, Oceanic Fm., Barbados]; [Not Cushman, 1927]
1974 Morozovelloides bandyi Fleisher. - Fleisher : p.1034 pl. 14; fig. 3-8 [Middle Zone P10 to Zone P1 1, DSDP Sites 219 and 220, Arabian Sea]
1993 Morozovella crassata Cushman. - Pearson et al. : pl. 2; fig. 4-6 [middle Eocene Zone P1 1-P1 2, DSDP Site 523, Walvis Ridge, south Atlantic Ocean]; [Not Cushman, 1925]
2006 Morozovelloides bandyi Fleisher. - Pearson et al. : p.333 pl. 10.1; fig. 1-16 (Pl. 10.1, Figs. 1-3: new SEMs of holotype of Morozovella bandyi Fleisher)
References:

Cushman,J.A. and Barksdale,J.D.. (1930):
Eocene foraminifera from Martinez, California . Stanford University, Department of Geology, Contributions Vol. 1 p. 55-73

Cushman,J.A. (1939):
Eocene Foraminifera. from submarine cores off the eastern coast of North America . Contr. Cushman Lab. Foram. Res. Vol. 15 p. 49-76

Beckmann,J.P. (1953):
Die Foraminiferen der Oceanic Formation (Eocaen-Oligocaen) von Barbados. KI. Antillen . Eclogae Geologicae Helvetiae Vol. 46 p. 301-412

Fleisher,R.L. (1974):
Cenozoic planktonic foraminifera and biostratigraphy, Arabian Sea, Deep Sea Drilling Project, Leg 23A.
In: Initial Reports of the Deep Sea Drilling Project Vol. 23 Eds: Whitmarsh, R.B.Weser, O.E.Ross, D.A. p. 1001-1072

Pearson,P.N.; Shackleton,N.J. and Hall,M.A. (1993):
The Stable Isotope Paleoecology of Middle Eocene Planktonic Foraminifera and Multi-species integrated Isotope Stratigraphy . Journal of Foraminiferal Research Vol. 23 p. 123-140

Pearson,P.N.; Olsson,R.K.; Hemleben,C.; Huber,B.T. and Berggren,W.A. (2006):
Atlas of Eocene Planktonic Foraminifera. p. 1-513
Anonymous: Unedited TaxonConcept data
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