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Species Hoploscaphites pungens-schmidi



Discussion / Comments:
Machalski (2005):
MATERIAL.- From Poland: a single specimen from Che³m (ZPAL Am. 12/706). From Denmark: a single specimen from Gudumholm, Jylland (MGUH 27745). From Germany: a single specimen from Hemmoor (NLfB kma 178). DISCUSSION.- According to Kennedy (1987: 208), his Acan− thoscaphites sp. “differs from the other Acanthoscaphites from the Limburg Maastrichtian in the flexuosity of the flank ribs, presence of only a single order of flank ribs (separating it from A. verneuilianus) and no umbilical bullae (separating it from H. pungens)”. The specimen described by Kennedy (1987) is from the Kunrade Limestone facies of the Maas− tricht Formation at Kunrade and consists of half of a whorl, septate for the largest portion (Kennedy 1987: pl. 32: 15–17). Tuberculation consists of distinct ventrolaterals and incipient laterals; no umbilical bullae are visible. Primary ribs are flexuous and prorsiradiate on the flanks; secondaries origi− nate at the junction of flank and ventrolateral shoulder. All specimens assigned herein share the rib flexuosity with Acanthoscaphites sp. of Kennedy (1987). Specimen ZPAL Am. 12/706 (Fig. 18C) comes from level III at Cheùm. It is a fragment of a spire plus the early part of the body chamber, 54 mmin maximum preserved diameter. Specimen ZPAL Am. 12/706 shares the presence of distinct ventro− lateral tubercles and incipient lateral tubercles with that from Kunrade. In addition, it has umbilical bullae, which are pres− ent in H. pungens and absent from the Kunrade individual. Specimen MGUH 27745 (Fig. 18B1, B2) is from the Gu− dumholm section, northern Jylland (Meonia semiglobularis– Ruegenella humboldtii Zone of Surlyk 1984: fig. 3). It is a fragment of phragmocone with part of the body chamber pre− served, 48 mm in maximum preserved diameter. The speci− men is probably a macroconch which can be inferred from a convexity of the preserved part of the umbilical wall of the shaft. The tuberculation in MGUH 27745 follows the pattern seen in the Kunrade specimen but the lateral tubercles streng− then markedly adaperturally. There are no umbilical bullae on the specimen, a feature shared with the Kunrade individual. Specimen NLfB kma 178 (Fig. 18A) is from 20.1 metres above the lower/upper Maastrichtian boundary clay M900 (T100) at Hemmoor, thus from the Sp. tegulatus–Bt. junior Zone of Schulz and Schmid (1983). It is a part of the phragmo− cone with the basal portion of the body chamber. An umbilical swelling is seen, which indicates this to be a macroconch. The original photograph of the specimen (Birkelund 1982: pl. 1: 4) is inadequate and the specimen is refigured herein. The latter shows ribbing similar to that in other specimens in the lot dis− cussed here: no umbilical bullae, incipient lateral tubercles and much stronger ventrolaterals. A specimen from Rügen illustrated by Wolansky (1932) differs from Hoploscaphites constrictus and shares a similar ornament with the present group, although the quality of the photograph does not allowto establish its tuberculation pattern. ?Acanthoscaphites sp. sensu Kennedy, 1987 in Jagt (1995) from the upper Nekum Member (Maastricht Formation) at the CBR−Romontbos quarry, Belgium, is represented by spires of a small diameter which may or may not belong here. In summary, the present specimens recall both H. pun− gens and H. schmidi in tuberculation, but differ in showing flexuous primary ribs on the phragmocone and early part of the body chamber. This material probably represents a new species of the H. pungens–schmidi group, but poor preserva− tion precludes a formal description. Alternatively, these spe− cimens could be end members of H. pungens and H. schmidi populations. STRATIGRAPHIC AND GEOGRAPHIC RANGE.- Upper lower Maas− trichtian and/or lower upper Maastrichtian of Denmark (Me− onia semiglobularis–Ruegenella humboldtii Zone), Germany (Spyridoceramus tegulatus–Belemnitella junior Zone), Po− land (Spyridoceramus tegulatus–Belemnitella junior Zone) and, possibly, of Belgium(Belemnitella junior Zone of authors).
Systematics:

51
 Familia Scaphitidae
  Genus Hoploscaphites
   Species Hoploscaphites pungens-schmidi
Synonym list:
Machalski (2005):
p 1932 Hoploscaphites constrictus Sowerby. - Wolansky : p.10 pl. 1; fig. 10 [non pl. 1, fig. 6 = Hoploscaphites tenuistriatus; non pl. 1, fig. 12 = Hoploscaphites constrictus]
p 1982 Acanthoscaphites varians Lopuski. - BIRKELUND : p.16 pl. 1; fig. 4 [non pl. 1, fig. 5 = Acanthoscaphites varians blaszkiewiczi; non pl. 1, fig. 6 = Hoploscaphites schmidi]
1987 Acanthoscaphites sp. . - Kennedy : p.208 pl. 32; fig. 15-17
? 1995 Acanthoscaphites? sp. sensu Kennedy. - Jagt : p.32 pl. 7; fig. 5, 6, 8, 9
2005 Hoploscaphites sp. ex gr. pungens-schmidi . - Machalski : p.678 fig. 18
Stratigraphy - relative ages:
upper Maastrichtian - lower Maastrichtian: Machalski (2005)
References:

Wolansky,D.. (1932):
Die Cephalopoden und Lamellibranchiaten der Ober-Kreide Pommerns . Geologie aus dem Geologisch-Paläontologischen Institut der Universität Greifswald Vol. 9 p. 72 pp.

BIRKELUND,T.. (1982):
Maastrichtian ammonites from Hemmoor, Niederelbe (NW Germany). . Geol. Jb. A61 p. 13-33

Kennedy,W.J.. (1987):
The ammonite faunas of the type Maastrichtian, with a revision of Ammonites colligatus Binkhorst, 1861 . Bulletin de l'Institut Royal des Sciences Naturelles de Belgique, Sciences de la Terre Vol. 56 p. 151-267

Jagt,J.W.M.. (1995):
A late Maastrichtian ammonite faunule in flint preservation from northeastern Belgium. . Mededelingen van de Rijks Geologische Dienst Vol. 53 p. 21-47

Machalski,M.. (2005):
Late Maastrichtian and earliest Danian scaphitid ammonites from central Europe: Taxonomy, evolution, and extinction . Acta Palaeontologica Polonica Vol. 50(4) p. 653-696

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