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Species Hoploscaphites felderi Kennedy 1987



Discussion / Comments:
Machalski (2005):
MATERIAL.- From the Netherlands: three specimens, one from quarry ‘t Rooth, Bemelen (NHMM MK 463), another from the ENCI quarry (NHMM JJ 11862a, b) and one from the former Blom quarry (NHMM JJ 7386a, b). From Bel− gium: two specimens from CBR−Romontbos quarry, Liège (a silicone castNHMM 1992074 of specimen no. 2015 in the Indeherberge collection and an unnumbered silicone cast of another specimen). From Poland: two specimens, one from Chełm chalk pit (ZPAL Am. 12/710) and one from the town quarry south of Kazimierz Dolny (ZPAL AM. 12/709). From Denmark: two specimens, one from “Danmark” quarry, Jyl− land (MGUH 20165), another a silicone cast of a specimen from Rørdal quarry, Ålborg (MGUH 27746). DISCUSSION.- No body chambers are yet known of this spe− cies, which is characterised by a compressed whorl section, flattened flanks, distinctive ornament and intricately subdi− vided suture (Kennedy 1987; Jagt 1995). Topotypes from the Kunrade Limestone facies (Kennedy 1987: pl. 33: 1–8, 10; pl. 34: 7–9, 12–17) do not differ in any respect of their mor− phology from the holotype. The same concerns other mate− rial from the Maastricht area, which is from the Emael and Nekum members of the Maastricht Formation (Jagt 1995: 31; Jagt 2002). Prior to the present study, Hoploscaphites felderi was known exclusively from Belgium and the Netherlands. New finds from Poland and a reinterpretation of some specimens from Denmark have now demonstrated a wider geographic range of this species. All this material consists of incomplete phragmocones, like the original material from the Maastricht area. This allows to speculate that the body chamber of H. felderi was either very thin or weakly calcified and thus prone to destruction shortly after the death of its inhabitant. Specimen ZPAL Am. 12/709 represents half a whorl, 33 mm in maximum preserved diameter (Fig. 19C). It comes from the lower part of the Kazimierz Opoka (Bn. kazimiro− viensis Zone) as exposed at the town quarry at Kazimierz Dolny, and shows distinctive ribbing, indistinguishable from that visible on external moulds of H. felderi from the Kunrade Limestone (Kennedy 1987: pl. 33: 2, 3) and on conspecific external moulds NHMM JJ 11862a, b (Fig. 19A) and NHMM JJ 7386a, b from the Nekum Member. In all these specimens, the primary ribs are concave on the inner flank, sweep forwards across mid−flank, and are concave on the outer flank. Secondaries arise on the inner flank, either by insertion or by subdivision of the primaries. In result, the outer flanks, ventrolateral shoulder and venter are covered by dense, sharp ribs. Another Polish specimen, ZPAL Am. 12/710 (Fig. 19D), is from level III at the Chełm chalk pit, from the Spyrido− ceramus tegulatus–Belemnitella junior Zone. It is an incom− plete spire, 25 mm in maximum preserved diameter, with the same ribbing pattern as described above and flattened flanks characteristic of the species. Specimen MGUH 20165 (Fig. 19E) from “Danmark” quarry, Jylland, is from the Meonia semiglobularis–Rue− genella humboldtii Zone (Surlyk 1984: fig. 3). It is 44 mm in maximum preserved diameter. The specimen is not H. tenui− striatus as claimed by Birkelund (1993: pl. 14: 14) as it is septate throughout and its ornament consists of fine, dense ribs on the ventrolateral shoulder and outer flank and rarer, stronger ribs on the mid− and inner flanks. Phragmocones of H. tenuistriatus tend to be smaller and are additionally cov− ered with rather coarse ribs. Specimen MGUH 20165 is closely similar in proportions and ribbing to an unnumbered silicone cast of H. felderi from the CBR−Romontbos quarry (Fig. 19F). In addition, MGUH 20165 has flattened flanks with a marked ventrolateral shoulder, typical of H. felderi. Another specimen from Denmark, MGUH 27746, is from Rørdal quarry, Ålborg, Jylland, from either the upper part of the Rugia tenuicostata–Meonia semiglobularis Zone or from the lower part of the Meonia semiglobularis–Ruege− nella humboldtii Zone (see Surlyk 1984: fig. 3). It is a con− torted fragment of a phragmocone whorl (Fig. 19B). Venter, ventrolateral shoulder and outer flank are visible, and are covered with sharp ribs typical of the species; only few pri− maries are visible on mid−flank, which, however, is poorly preserved. Parts of a deeply incised suture line are also seen. The size of MGUH 27746 suggests a much larger individual than the majority of other specimens of the species known to date. However, fragments NHMM JJ 11862a, b (Fig. 19A) and NHMM JJ 7386a, b from the Nekum Member, must also have belonged to fairly large individuals. Perhaps, two spe− cies or dimorphs are involved. This cannot be clarified until better−preserved specimens with body chambers are found. STRATIGRAPHIC AND GEOGRAPHIC RANGE.- Upper lower and/or lower upper Maastrichtian of Belgium and the Netherlands (Belemnitella junior Zone of authors), Denmark (Meonia semiglobularis–Ruegenella humboldtii Zone and possibly Rugia tenuicostata–Meonia semiglobularis Zone), and Po− land (Spyridoceramus tegulatus–Belemnitella junior Zone and Belemnella kazimiroviensis Zone).
Systematics:

51
 Familia Scaphitidae
  Genus Hoploscaphites
   Species Hoploscaphites felderi
Synonym list:
Machalski (2005):
1861 Ammonites n sp. Decheni Binkhorst. - Binkhorst : p.30 pl. 5a; fig. 15a-e
1908 Scaphites cf. roemeri d'Orbigny. - Grossouvre : p.35 pl. 10; fig. 1-3
1987 Hoploscaphites n sp. felderi Kennedy. - Kennedy : p.203 pl. 27, fig. 1; pl. 33, fig. 1-15; pl. 34, fig. 7-11, 13-17; text-fig. 13c
p 1993 Hoploscaphites tenuistriatus Kner. - BIRKELUND : p.59 pl. 14; fig. 4 [non pl. 14, fig. 8-11, 15, 16 = Hoploscaphites tenuistriatus; non pl. 14, fig. 13 = Hoploscaphites sp. ]
1995 Hoploscaphites felderi Kennedy. - Jagt : p.30 pl. 6, fig. 3-6; pl. 7, fig. 3, 4, 10-12
2005 Hoploscaphites felderi Kennedy. - Machalski : p.680 fig. 19
Stratigraphy - relative ages:
upper Maastrichtian - lower Maastrichtian: Machalski (2005)
References:

Binkhorst,J.T.. (1861):
Monographie des gastropodes et des céphalopodes de la Craie Supérieure du Limburg.
In: Maastricht and Muller Frères Eds: Muquardt, G.. p. 17,83,44

Grossouvre,d.A.. (1908):
Description des ammonites du Cretace Superieur du Limbourg Belge et Hollandais et du Hainault . Mem. Mus. r. Hist. nat. Belg Vol. 4 p. 1-39

Kennedy,W.J.. (1987):
The ammonite faunas of the type Maastrichtian, with a revision of Ammonites colligatus Binkhorst, 1861 . Bulletin de l'Institut Royal des Sciences Naturelles de Belgique, Sciences de la Terre Vol. 56 p. 151-267

BIRKELUND,T.. (1993):
Ammonites from the Maastrichtian White Chalk in Denmark . Bulletin of the Geological Society of Denmark Vol. 40 p. 33-81

Jagt,J.W.M.. (1995):
A late Maastrichtian ammonite faunule in flint preservation from northeastern Belgium. . Mededelingen van de Rijks Geologische Dienst Vol. 53 p. 21-47

Machalski,M.. (2005):
Late Maastrichtian and earliest Danian scaphitid ammonites from central Europe: Taxonomy, evolution, and extinction . Acta Palaeontologica Polonica Vol. 50(4) p. 653-696

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