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Subspecies Acanthoscaphites (Euroscaphites) varians varians Lopuski 1911



Alternative name:
Acanthoscaphites varians varians Lopuski 1911
Discussion / Comments:
Machalski (2005):
TYPE MATERIAL.- Holotype, by monotypy, is the original of Łopuski (1911: pl. 4: 1–3) from the upper upper Maastrichtian at Kazimierz Dolny, Middle Vistula River section, Poland. The specimen is presum− ably lost. MATERIAL.- From Poland: five specimens from the town quarry south of Kazimierz Dolny (ZPAL Am. 12/400, 778; MKD/MP−135; unregistered specimen in Marcinowski col− lection, unregistered specimen in the Praszkier and Dembicz collection) and six specimens from the Rejowiec quarry (ZPAL Am. 12/401, 776, 777, 779, 1279, unregistered speci− men in the Praszkier and Dembicz collection). DISCUSSION.- The specimens assigned herein to A. (E.) v. varians were treated in detail by Łopuski (1911), Błaszkie− wicz (1980), Machalski (1996), and Jagt et al. (1999) and there is no need to repeat these descriptions. Newly collected specimens from Rejowiec and Kazimierz Dolny (Fig. 21) match in their ribbing and tuberculation the previously re− corded material. All material studied comes from the lower part of the Belemnella kazimiroviensis Zone as exposed at the town quarry south of Kazimierz Dolny and at Rejowiec. The new finds indicate that A. (E.) v. varians is not restricted to certain parts of these sections as assumed by Jagt et al. (1999), but occurs throughout. No records of the subspecies outside Po− land are known to date. Both the holotype and other specimens known are best in− terpreted as macroconchs, being similar in size and shell pro− portions to those of Acanthoscaphites tridens (Kner, 1848) as described by Kennedy and Summesberger (1987) and Jagt et al. (1999). The largest macroconch of A. (E.) v. varians known to date is ZPAL Am. 12/401, a complete specimen with markedly constricted aperture and a maximum diameter of 256 mm (Jagt et al. 1999: fig. 3, pl. 6). This specimen co− mes from an opoka unit between the baculite and belemnite marls at Rejowiec. No specimens which could be interpreted as microconchs of A. (E.) v. varians are known yet. Specimen ZPAL Am. 12/152 from the town quarry south of Kazimierz Dolny was interpreted as microconch by Jagt et al. (1999). However, it has a smooth venter with indistinct siphonal swellings and is better regarded as unrelated to the present subspecies. STRATIGRAPHIC AND GEOGRAPHIC RANGE.- Upper upper Maas− trichtian of Poland (Belemnella kazimiroviensis Zone).
Systematics:

51
 Species Acanthoscaphites (Euroscaphites)
  Subspecies Acanthoscaphites (Euroscaphites) varians varians
Synonym list:
Machalski (2005):
1911 Scaphites n sp. varians Lopuski. - Lopuski : p.120 pl. 4; fig. 1-3
1980 Acanthoscaphites varians Lopuski. - BLASZKIEWICZ : p.40 pl. 25; fig. 1, 2, 5, 6
1996 Acanthoscaphites varians Lopuski. - Machalski : p.378 fig. 6
p 1999 Acanthoscaphites (Euroscaphites) varians varians Lopuski. - Jagt et al. : p.137 text-fig. 3, 4; pl. 3; pl. 4, fig. 2, 3; pl. 5; fig. 1-4; pl. 6; pl. 7 [non pl. 4; fig. 1]
non 2002 Acanthoscaphites varians Lopuski. - Reich & Frenzel : pl. 23; fig. 1a, b [= Acanthoscaphites (Euroscaphites?) sp. aff. verneuilianus]
2005 Acanthoscaphites (Euroscaphites) varians varians Lopuski. - Machalski : p. 683, 684 fig. 21, 22B
Stratigraphy - relative ages:
upper Maastrichtian: Machalski (2005)
References:

Lopuski,C.. (1911):
Przyczynld do znajomosci fauny kredowej gub. Lubelskiej. . Cr. Seanc. Soc. Sci. Varsovie Vol. 4 p. 104-140

BLASZKIEWICZ,A.. (1980):
Campanian and Maastrichtian ammmonites of the Middle Vistula Valley, Poland: a stratigraphic-paleontological study . Prace Instytutu Geologicznego Vol. 92 p. 1-63

Machalski,M.. (1996):
Scaphitid ammonite correlation of the Late Maastrichtian deposits in Poland and Denmark . Acta Palaeontologica Polonica Vol. 41 p. 369-383

Jagt,W.J.M..; Kennedy,W.J.. and Machalski,M.. (1999):
Giant scaphitid ammonites from the Maastrichtian of Europe . Bulletin de l’Institut royal des Sciences naturelles de Belgique, Sciences de la Terre Vol. 69 p. 133-154

Reich,M.. and Frenzel,P.. (2002):
Die Fauna und Flora der Rügener Schreib− kreide (Maastrichtium, Ostsee) . Archiv für Geschiebekunde Vol. 3 p. 74-284

Machalski,M.. (2005):
Late Maastrichtian and earliest Danian scaphitid ammonites from central Europe: Taxonomy, evolution, and extinction . Acta Palaeontologica Polonica Vol. 50(4) p. 653-696

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