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Species Nucula ornata Sowerby 1846



Discussion / Comments:
Griffin & Nielsen (2008):
TYPE MATERIAL. Lectotype (designation by del Río & Camacho 1998): NHM-L27974; paralectotype: NHM-L27975 REMARKS. Sowerby (p. 251) noted that “a fragment alone has been found”. However, in the collection there are two specimens: one of them is a fragment, but the other one (NHM-L27974) is the right valve figured by Sowerby (1846, pl. 2, fig. 19). This specimen is a broken right valve attached to the internal mould, and in which the anterior end is missing. It seems to have been repaired after being illustrated by Sowerby, as the crack is clearly visible, and all other features indicate it is obviously the same specimen. It remains unclear where the second fragment (NHM-L27975) comes from. The fact that Sowerby mentions “only a fragment” may have led del Río & Camacho (1998) to believe it was the holotype, but there are in fact two syntypes. Del Río & Camacho (1998) stated that “the holotype is a broken right valve found in the lower fossiliferous strata that crop out at Puerto Deseado (Santa Cruz Province)”. It is not clear where they drew this conclusion from, as Darwin did not mention what beds his material came from. Nevertheless, it seems likely that they are right. The fossil beds exposed near Puerto Deseado (a few kilometers inland) are probably equivalent to the late Oligocene-early Miocene Monte León Formation. In any event, there was no way of knowing or inferring from the original description that there was more than one specimen and therefore their lectotype designation by inference of holotype is valid (ICZN 1999, Art. 74.6).
Synonym list:
Griffin & Nielsen (2008):
1846 Nucula ornata Sowerby. - Sowerby : p.251 pl. 2; fig. 19
1899 Neilo ornata Sowerby. - Ihering : p.14
1907 Malletia ornata Sowerby. - Ihering : p.228
1998 Neilo (Neilo) ornata Sowerby. - del Rio & Camacho : 65-67 pl. 2; fig. 11-14; text-fig. 7
1999 Neilo ornata Sowerby. - Frassinetti & Covacevich : p.16 pl. 2; fig. 10-12
2008 Neilo ornata Sowerby. - Griffin & Nielsen : pl. 8; fig. 8-9
References:

Sowerby,G.B.. (1846):
Descriptions of Tertiary fossil shells from South America..
In: Geological observations on South America. Eds: Darwin, C.. p. 249-264

Ihering,v.H.. (1899):
Conchylien der patagonischen Formation . Neues Jahrbuch für Mineralogie, Geologie und Palaontologie p. 1-46

Ihering,H.. (1907):
Les mollusques fossiles du Tertiaire et du Crétacé Supérieur de l’Argentine . Anales del Museo Nacional de Buenos Aires 3/7 p. 611

del Rio,C.J.. and Camacho,H.H.. (1998):
Tertiary nuculoids and arcoids of eastern Patagonia (Argentina) . Palaeontographica A Vol. 250 p. 47-88

Frassinetti,D.. and Covacevich,V.. (1999):
Invertebrados fosiles marinos de la Formacion Guadal (Oligoceno superior-Mioceneo inferior) en Pampa Castillo, region de Aisen, Chile Translated Title: Invertebrate marine fossils from the Guadal Formation (upper Oligocene-lower Miocene) at Pampa Castillo, Aisen region, Chile.. Vol. 51 p. 96

Griffin,M.. and Nielsen,S.N.. (2008):
A revision of the type specimens of Tertiary molluscs from Chile and Argentina described by d’Orbigny (1842), Sowerby (1846), and Hupé (1854) . Journal of Systematic Palaeontology

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