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Species Trochus collaris Sowerby 1846

Discussion / Comments:
Griffin & Nielsen (2008):
TYPE MATERIAL. There is one syntype in the NHM under number NHM-G26397. The other material referred to this species by Sowerby could not be located in the Darwin Collection at the NHM. As discussed below, the lost specimen from Navidad was probably a juvenile of Astele chilensis (d’Orbigny, 1852). In order to fix the status of this species we designate specimen NHM-G26397 as lectotype of Valdesia collaris (Sowerby, 1846). REMARKS. The first species referable to Valdesia described from the Patagonian Tertiary was Trochus collaris Sowerby, 1846 (p. 613, pl. 3, figs 44-45). This species was reported by Sowerby from Navidad and Santa Cruz. The specimen (NHM-G26937) he figured in pl. 3, fig. 45 reads Navidad on the label. However, according to handwriting on the original label by Martin Doello Jurado it comes from Santa Cruz, a locality also confirmed on the same label by W.J. Zinsmeister. We agree with the opinion that this specimen comes from Santa Cruz and not from Navidad. Sowerby’s specimen from Navidad seems lost and appears to have been a juvenile of Astele chilensis, as already suspected by Sowerby himself who believed it might have been a juvenile stage of Trochus laevis, which he collected only in Navidad (Chile). Philippi (1887) agreed with Sowerby on this issue, and synonymised the two species, which were later placed under Gibbula by Tavera (1979). However, the shells from Navidad and Santa Cruz are in fact quite different and can not be placed under the same specific name. The specimens from Santa Cruz have an almost smooth shell with only a row of faint tubercles immediately below the suture and very faint spiral cords on the early teleoconch whorls. The periphery of the shell is bicarinate, visible as such only on the last whorl. The base has about 10 spiral cords, of which the three or four nearest to the umbilicus are the strongest. Besides, the whorls in Astele chilensis (= Trochus laevis Sow.) are more weakly inflated than in Valdesia collaris. Until all the fossil material of these species is better known, it is preferable to use Valdesia collaris for the material from Santa Cruz and Astele chilensis for the material from Navidad.
Synonym list:
Griffin & Nielsen (2008):
1846 Trochus collaris Sowerby. - Sowerby : p.256 pl. 3; fig. 44, 45
1902 Gibbula laevis Sowerby. - Ortmann : p.170 pl. 31; fig. 8
1987 Valdesia (Juliania) collaris Sowerby. - Morra & del Rio : p.87 pl. 2; fig. 3a-c
1999 Trochus laevis Sowerby. - Frassinetti & Covacevich : p.45 pl. 9; fig. 5-9
2008 Valdesia collaris Sowerby. - Griffin & Nielsen : pl. 20; fig. 10-11
Was used in synonym list of:
Astele chilensis d'Orbigny 1852
Trochus laevis Sowerby 1846

Sowerby,G.B.. (1846):
Descriptions of Tertiary fossil shells from South America..
In: Geological observations on South America. Eds: Darwin, C.. p. 249-264

Ortmann,A.E.. (1902):
Tertiary invertebrates: Reports of the Princeton Expedition to Patagonia, 1896–1899. Vol. 4(2) p. 45-332

Morra,G.A.. and del Rio,C.J.. (1987):
. La subfamilia Architectonicinae (Mollusca, Gastropoda) en el “Patagoniano” de la costa atlántica, Chubut y Santa Cruz . Revista de la Asociación Geológica Argentina Vol. 42 p. 82-91

Frassinetti,D.. and Covacevich,V.. (1999):
Invertebrados fosiles marinos de la Formacion Guadal (Oligoceno superior-Mioceneo inferior) en Pampa Castillo, region de Aisen, Chile Translated Title: Invertebrate marine fossils from the Guadal Formation (upper Oligocene-lower Miocene) at Pampa Castillo, Aisen region, Chile.. Vol. 51 p. 96

Griffin,M.. and Nielsen,S.N.. (2008):
A revision of the type specimens of Tertiary molluscs from Chile and Argentina described by d’Orbigny (1842), Sowerby (1846), and Hupé (1854) . Journal of Systematic Palaeontology

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