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Species Hauericeras rembda Forbes 1846

[1]
[2]
[1] pl. 2, figs. 4-6 Ifrim et al. (2004) [2] text-fg. 8e-f Ifrim et al. (2004)

Alternative name:
Hauericeras (Gardeniceras) rembda Forbes 1846
Diagnosis / Definition:
Cobban & Kennedy (1995):
DESCRIPTION.- USNM 463101, a wholly septate phosphatic fragment 30 mm long, has compressed whorl section and whorl breadth to height ratio of 0.61. Coiling evolute, with shallow umbilicus, low, feebly convex and subvertical umbilical wall, and narrowly rounded umbilical shoulder. Flanks slightly convex and subparallel with greatest breadth around midflank. Venter narrowly rounded with traces of what appears to be floor of a keel in places. Surface of shell smooth, but for single prominent, broad, markedly biconcave, prorsiradiate constriction projected strongly forward and weakened on venter to form narrowly rounded ventral chevron. Suture has large ventral saddle, narrow-stemmed E/L, and large, bifid, deeply incised L, and bifid L/U, as large as E/L and similarly incised; U, smaller and asymmetrically bifid, remainder of U being strongly retracted.
Ifrim et al. (2004):
Description: Evolute with low to intermediate expansion rate. The flat umbilicus spans c. 40 per cent of the shell diameter. Approximately one-third of the previous whorl is covered. The flanks are subparallel and slightly convex, with a short vertical wall and narrowly rounded umbilical shoulders. Maximum whorl breadth is at the middle flank. WB/WH decreases with growth due to a gradually compressed whorl section. Three to four deep biconvex constrictions are present per whorl. They are convex on the middle flank and concave and prorsiradiate on the outside to cross the venter in a projected bow. From a diameter of 33 mm onwards a faint keel is preserved on the largest specimen (UANL CE MAAS-063). E is broad and much shallower than the asymmetric trifid L. The umbilical suture line is slightly (early growth stages) to strongly (later growth stages) retracted.
Ward & Kennedy (1993):
Description. - Coiling very evolute with umbilicus comprising up to 37 percent of diameter, with low umbilical wall and nar- rowly rounded shoulder. Whorls expand slowly; composite molds smooth, but for distant narrow constrictions, four or five per whorl. These are feebly prorsiradiate, concave on inner flank, convex at mid-flank, and concave on outer flank, projecting strongly forwards on ventrolateral shoulder. Strong siphonal keel. Sutures not seen.
Discussion / Comments:
Cobban & Kennedy (1995):
TYPES.- The lectotype is BMNH C5 1024, the original of Forbes, 1846, plate 7, figure 3, by the subsequent designation of Matsumoto and Obata, 1955, page 145; paralectotypes are BMNH C5 1023 and '251025, from the Valudavur Formation of Pondicheny, South India. Material.- A single specimen, USNM 463101. DISCUSSION.- The markedly biconcave constrictions distinguish Hauericeras rembda from all other species referred to the genus. A full revision of this species and a detailed discussion of differences from its allies is given by Kennedy and Henderson (1992a), to which the reader is referred. OCCURRENCE.- Upper Maastrichtian of South India and the Biscay region of Spain; Maastrichtian of Madagascar and Zululand (South Africa). The present specimen is from the Prairie Bluff Chalk at locality 34.
Ifrim et al. (2004):
Remarks: Hauericeras rembda is more involute than H.fayoli (de Grossouvre, 1894) (for comparison, see Kennedy 1986c). Kennedy and Henderson (l992b) showed that H. rembda has a keel from a diameter of 20 mm onwards. In our material the keel appears only on our largest specimen at a diameter of 33 mm. The Cerralvo specimens displayajuvenile suture line which is less complex even for the largest specimen than the suture line described by Kennedy and Henderson (l992b). Occurrence: Hauericeras rembda (Forbes) is known from Japan (Matsumoto and Obata 1955), tbe Gansserina gansseri Zone in tbe Biscay region (Ward and Kennedy 1993), tbe Upper Maastrichtian of the Valudavur Formation in soutbern India (Kennedy and Henderson 1992b), and the Maastrichtian of Madagascar (Collignon 1971). A single fragment has been recorded from tbe Maastrichtian of Alabama (Cobban and Kennedy 1995).
Ward & Kennedy (1993):
Types.- Lectotype, by the subsequent designation of Matsumoto and Obata (1 955, p. 145), is BMNH C5 1024, the original of Forbes 1846 (Pl. 7, fig. 3), paralectotypes are BMNH C5 1023 and C5 1025. The figured syntype of Ammonites Durga Forbes, 1846 (p. 104, Pl. 7, fig. 1 1) is BMNH C5 102 1, all from the Valudavur Formation of Pondicheny, south India (ex Kaye and Cunliffe Collection). BMNH (23567 is a topotype. Discussion. - As noted by Kossmat (1 898), Ammonites Durga of Forbes represents the early growth stages of Hauericeras rembda, prior to development of the keel. This is well illustrated by the largest paralectotype, BMNH C51023, which shows the beginning of a keel developing at 20 mm. The syntypes of durga, BMNH C5 1021 and 5 1022, are both less than 20 mm diameter. Hauericeras rembda most closely resembles Hauericeras fayoli de Grossouvre, 1894 (p. 220 (pars), Pl. 27, fig. 3), which is probably from Tercis, Landes, France, and probably of late Campanian age (see revisions in Kennedy and Summesberger, 1984, p. 157, Pl. 1, figs. 8, 12, Pl. 2 figs. 4, 13, 5, and Kennedy, 1986a, p. 27, text-fig. 9a-j). But this species is much more evolute, with U = 45 percent, and constrictions that are sinuous rather than biconvex. The Maastrichtian H. sulcatum (Kner, 1848) (p. 8, Pl. 1, fig. 3; see revision in Kennedy and Summesberger, 1987, p. 27, Pl. 1 figs. 1-7, Pl. 13, fig. 2) has 6-7 prorsiradiate constrictions per whorl that are concave, rather than biconvex. Occurrence.- Maastrichtian of south India; in the Biscay sections this taxon is rare and short ranging; it has been found only over a narrow stratigraphic interval near the top of Member I (lower Maastrichtian).
Synonym list:
Cobban & Kennedy (1995):
1992 Ammonites rembda Forbes. - Kennedy & Henderson : p.408 pl. 6, fig. 10-24; pl. 17, fig. 1; text-fig. 3H [with full synonymy]
1995 Hauericeras rembda Forbes. - Cobban & Kennedy : p. 5, 6 fig. 2.6, 2.7, 3
Ifrim et al. (2004):
1846 Ammonites rembda Forbes. - Forbes : p.111 pl. 7, fig. 3
1846 Ammonites durga Forbes. - Forbes : p.104 pl. 7, fig. 11
1898 Desmoceras (Hauericeras) rembda Forbes. - Kossmat : 124 (189) pl. 18, fig. 9
1955 Hauericeras (Gardeniceras) cf. rembda Forbes. - Matsumoto & Obata : p.144 pl. 29, figs. 6-7; text-fig. 13
1971 Hauericeras (Gardeniceras) rembda Forbes. - COLLIGNON : p.37 pl. 655, fig. 2417
1992 Hauericeras rembda Forbes. - Kennedy & Henderson : p.408 pl. 6, figs. 10-24; pl. 17, fig. 1; text-fig. 3h (with full synonymy)
1993 Hauericeras rembda Forbes. - Ward & Kennedy : p.24 fig. 16.7
1995 Hauericeras rembda Forbes. - Cobban & Kennedy : p.4 figs. 2.6, 2.7, 3
2004 Hauericeras rembda Forbes. - Ifrim et al. : 23, 20 pl. 2, figs. 4-6; text-fig. 8e-f
Ward & Kennedy (1993):
1846 Ammonites rembda Forbes. - Forbes : p.111 pl. 7; fig. 3
1846 Ammonites durga Forbes. - Forbes : p.104 pl. 7; fig. 11
p 1850 Ammonites rembda Forbes. - d'Orbigny : p.213
p 1850 Ammonites durga Forbes. - d'Orbigny : p.213
1865 Ammonites rembda Forbes. - Stoliczka : p.63 pl. 33; fig. 5
1865 Ammonites durga Forbes. - Stoliczka : p.143 pl. 7; fig. 5 only [non fig. 6, 7; =Puzosia compressa Kossmatt, 1898]
non 1871 Ammonites rembda Forbes. - GRIESBACH : p.63 pl. 3; fig. 2, 3 [=Hauericeras gardeni Baily, 1855]
1898 Desmoceras (Hauericeras) sp. rembda Forbes. - Kossmat : 124 (198) pl. 18(24); fig. 9
non 1906 Ammonites rembda Forbes. - WOODS : p.333 [=Hauericeras gardeni]
non 1907 Hauericeras rembda Forbes. - PERVINQUIERE : p.167 pl. 7, fig. 7-10; text-fig. 69 [=Hauericeras fayoli de Grossouvre, 1894]
1955 Hauericeras (Gardeniceras) cf. rembda Forbes. - Matsumoto & Obata : p.144 pl. 29; fig. 6, 7
1955 Hauericeras rembda Forbes. - Matsumoto & Obata : p.145 text-fig. 13
1971 Hauericeras rembda Forbes. - COLLIGNON : p.37 pl. 655; fig. 2417
1977 Hauericeras rembda Forbes. - Kennedy : text-fig. 31, 18
1993 Hauericeras rembda Forbes. - Ward & Kennedy : p.16 fig. 16.7
Stratigraphy - relative ages:
Maastrichtian: Ifrim et al. (2004)
lower Maastrichtian - lower Maastrichtian: Ward & Kennedy (1993)
References:

Forbes,E.. (1846):
Report on the Cretaceous fossil invertebrates from southern India, collected by Mr. Kaye and Mr. Cunliffe . Transactions of the Geological Society of London Vol. 2(7) p. 97–174

d'Orbigny,A. (1850):
Prodrome de Paléontologie. Stratigraphique universelle des animaux mollusques & rayonnés faisant suitre au cours élémentaire de paléontologie et de géologie stratigraphique. , Cours Élémentaire de Paléontologie et de Géologie Stratigraphiques Vol. 2

Stoliczka,F.. (1865):
The fossil Cephalopoda of the Cretaceous rocks of southern India. Ammonitidae with revision of the Nautilidae etc. . Memoirs of the Geological Survey of India, (1), Palaeontologica Indica6-9 p. 107-154

GRIESBACH,C.L.. (1871):
On the geology of Natal in South Africa . Quarterly Journal ofthe Geological Society of London Vol. 27 p. 53-72

Kossmat,F.. (1898):
Untersuchungen über die Südindische Kreideformation. Dritter Theil. . Beiträge zur Paläontologie Österreich-Ungarns und des Orient Vol. 11 p. 89-152

WOODS,H.. (1906):
The Cretaceous fauna of Pondoland . Annals of the South African Museum Vol. 4 p. 275-350

PERVINQUIERE,L.. (1907):
Etudes de paleontologie tunisienne I: Cephalopodes des terrains secondaires.
In: Carte Geologique de Tunisie p. 438 pp

Matsumoto,T.. and Obata,I. (1955):
Some Upper Cretaceous desmoceratids from Hokkaido and Saghalien . Memoirs of the Faculty of Science, Kyushu University, Series D, Geology Vol. 5 p. 119-151

COLLIGNON,M.. (1971):
Atlas des fossiles caracteristiques de Madagascar (Ammonites)(Maestrichtian). Vol. 17 p. 82

Kennedy,W.J.. (1977):
Ammonite evolution.
In: Patterns of Evolution Eds: Hallam, A.. p. 251-330

Kennedy,W.J.. and Henderson,R.A.. (1992):
Non-heteromorph ammonites from the Upper Maastrichtian of Pondicherry, south India . Palaeontology Vol. 35 p. 381-442

Kennedy,W.J.. and Henderson,R.A.. (1992):
Maastrichtian ammonites (Phylloceratina, Lytoceratina, Ammonitina) from Pondicheny, South India . Palaeontology Vol. 35 p. 381-442

Ward,P.D.. and Kennedy,W.J.. (1993):
Maastrichtian Ammonites from the Biscay Region (France, Spain) . Memoir (The Paleontological Society), Journal of Paleontology34 (Supplement to Vol. 67)(5) p. 1-58

Ward,P.D.. and Kennedy,W.J.. (1993):
Maastrichtian ammonites from the Biscay Region . Journal of Paleontology Vol. 67(5 II) p. 1-58

Cobban,W.A.. and Kennedy,W.J.. (1995):
Maastrichtian ammonites chiefly from the Prairie Bluff Chalk in Alabama and Mississippi . Journal of Paleontology Vol. 69(5) p. 1-40

Ifrim,C..; Stinnesbeck,W.. and López-Oliva,J.G.. (2004):
Maastrichtian cephalopods from Cerralvo, north-eastern Mexico . Palaeontology Vol. 47(6) p. 1575–1627

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