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Species Turborotalia altispiroides Bermudez 1961



Diagnosis / Definition:
Pearson et al. (2006):
DESCRIPTION. Type of wall: Smooth with pustules or weakly cancellate, especially on earlier chambers; tendency to defoliate. Test morphology: Highly trochospiral, globular, spherical morphology with 4 1/2 to 6 chambers in the final whorl and up to 3 1/2 whorls in the test; chambers moderately inflated, radially compressed but elongate in a dorso-ventral direction, increasing slowly in size, with final one or two chambers commonly flattened, reduced in size and added high in the spiral and arching over umbilicus; dorsal sutures slightly curved, depressed. Aperture a broad arch overhanging umbilicus, with narrow imperforate lip; umbilicus varying from narrow in specimens with 4 1/2 chambers in the final whorl to broad and deep in specimens with more; ventral sutures slightly curved, depressed; weak tendency for sinistral coiling. Size: Holotype length 0.51 mm, breadth 0.42 mm, height 0.46 mm.
Discussion / Comments:
Pearson et al. (2006):
DISTINGUISHING FEATURES.- Turborotalia altispiroides is distinguished from T pomeroli by its - higher spire, more dorso-ventrally extended chambers, and usually by having more chambers in the final whorl and a more umbilically-centered aperture. DISCUSSION.- This taxon has rarely been identified, despite being a distinctive and stratigraphically useful form. We illustrate the holotype in SEM for the first time (Pl. 15.1, Figs. 1-3). It is a large, high-spired species that, as originally noted by Bermudez (1961), is reminiscent in general shape of Neogene Dentoglobigerina altispira( Cushman and Jarvis) although it lacks the distinct umbilical teeth of that taxon. We have observed extreme variants in ODP Site 865 (equatorial mid Pacific Ocean - see P1. 15.1, Fig. 11). Morphometric studies (P.N. Pearson, unpublished data) show that it is a separate species from T. pomeroli, although the two intergrade in the lower part of the range of altispiroides. Its extinction in Zone E14 may prove to be a useful marker horizon, although it needs further documentation. PHYLOGENETIC RELATIONSHIPS.- Evolved from Turborotalia pomeroli and became extinct in the middle Eocene. STRATIGRAPHIC RANGE.- Middle Eocene, upper Zone Ell to lower Zone E14. GEOGRAPHIC DISTRIBUTION.- Probably cosmopolitan. STABLE ISOTOPE PALEOBIOLOGY.- Oxygen and carbon isotopic values are similar to T. pomeroli and intermediate in comparison to other species (P.N. Pearson, unpublished data).
Systematics:

35
 Ordo Foraminiferida
  Superfamilia Globigerinaceae
   Familia Hedbergellidae
    Genus Turborotalia
     Species Turborotalia altispiroides
Synonym list:
Pearson et al. (2006):
1953 Globorotalia centralis Cushman & Bermudez. - Beckmann : p.396 pl. 26; fig. 8-9 [middle-upper Eocene, Oceanic Fm., Barbados]
p 1957 Globorotalia centralis Cushman & Bermudez. - Bolli : p.169 pl. 39; fig. 2a-b [middle Eocene Porticulasphaera mexicana Zone. Navet Fm., Trinidad]; [Not Cushman and Bermudez, 1937]
1961 Turborotalia altispiroides Bermudez. - Bermudez : p. 1316-1617 pl. 17; fig. 4a-b [upper Eocene Jacabo Fm., Cuba]
2006 Turborotalia altispiroides Bermudez. - Pearson et al. : p.439 pl. 15.1; fig. 1-15 (Pl. 15.l, Figs. 1-3; new SEMs of holotype of Turborotalia altispiroides Bermudez)
Specimen:
Smithsonian Museum of Natural History, Washington, D.C., Inventory number: USNM 639071
References:

Beckmann,J.P. (1953):
Die Foraminiferen der Oceanic Formation (Eocaen-Oligocaen) von Barbados. KI. Antillen . Eclogae Geologicae Helvetiae Vol. 46 p. 301-412

Bolli,H.M. (1957):
Planktonic Foraminifera from the Eocene Navet and San Fernando formations of Trinidad, B.W.I. . Bull. U.S. natl. Mus. Vol. 215 p. 155-172

Bermudez,P.J. (1961):
Contribucion al estudio de las Globigerinidea de la region Caribe-Antillana (Paleocene-Reciente).
In: Boletino Geologia (Venezuela), Special Publicacion Vol. 3

Pearson,P.N.; Olsson,R.K.; Hemleben,C.; Huber,B.T. and Berggren,W.A. (2006):
Atlas of Eocene Planktonic Foraminifera. p. 1-513

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