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Species Hantkenina nuttalli Toumarkine 1981



Discussion / Comments:
Toumarkine & Luterbacher (1985):
Hantkenina nuttalli, the earliest representative of the genus Hantkenina, differs from the other species of the genus by its large size (0.5 to 1.0 mm) and the stellate shape of the test. The chambers (4 to 5) of the last whorl increase rapidly in size and are well separated from each other and radially elongate. Hantkenina mexicana Cushman is very close to H. nuttalli, but it is generally smaller and has longer spines. The general shape of the test is not as stellate as in H. nuttalli and the chambers become slightly imbricated. H. lehneri Cushman & Jarvis (Fig. 23.7) has more (5 to 7) chambers in the last whorl and these are more cylindrically finger-shaped. Nuttall (1930) described a variety of H. mexicana, H. mexicana var. aragonensis, but he illustrated three specimens without designating a holotype. Thalmann (1942b) raised the variety to specific rank and designated the specimen figured by Nuttall on pi. 24, fig. 1 as lectotype. Bolli et al. (1957) erroneously re-illustrated the specimen of fig. 3 of Nuttall (1930) instead of the one of his fig. 1 as the lectotype of H. aragonensis. Ramsay (1962) noticed this mistake and remarked that the real lectotype of H. aragonensis (pi. 24, fig. 1 of Nuttall, 1930) is very close to the holotype of H. mexicana. He, therefore, placed H. aragonensis into the synonymy of H. mexicana. His conclusions were also followed by Blow (1979). Toumarkine (1981) has come partly to the same conclusions: the holotype of H. aragonensis is a junior synonym of H. mexicana, but the paratype re-illustrated by Bolli et al. (1957, pi. 2, figs. 3a-b = Nuttall, 1930, pi. 24, fig. 3) is considered as a separate taxon which she names H. nuttalli.
Systematics:

1
 Superregnum Eukaryota
  Regnum Protoctista
   Phylum Ciliophora
    Subphylum Postciliodesmatophora
     Ordo Globigerinida
      Superfamilia Hantkeninacea
       Superfamilia Nonionacea
        Familia Hantkeninidae
         Genus Hantkenina
          Species Hantkenina nuttalli

15
  Classis Foraminifera
   Genus Hantkenina
    Species Hantkenina alabamensis
     Species Hantkenina nuttalli
Synonym list:
Toumarkine & Luterbacher (1985):
1930 Hantkenina mexicana var. aragonensis Nuttall. - Nuttall : pl 24 fig 3
1957 Hantkenina mexicana var. aragonensis Nuttall. - Bolli et al. : pl 2 figs 3a-b
1981 Hantkenina nuttalli Toumarkine. - Toumarkine : p.112 pl 1 fig 4, 5 and 6 (fig 4 from Nuttall, 1930, pl 24 fig 3 paratype of H. mexicana var. aragonensis; fig 5 and 6 from Nuttall, 1930 redrawn in Bolli at al 1957)
1985 Hantkenina nuttalli Toumarkine. - Toumarkine & Luterbacher : p.122 figs 23.1-5
Was used in synonym list of:
Hantkenina mexicana Cushman 1924
Stratigraphy - absolute ages:
FAD: 49 ± 0 [Ma], Berggren et al. (1995)
References:

Nuttall,W.L.F. (1930):
Eocene foraminifera from Mexico . Journal of Paleontology Vol. 4 p. 271-293

Thalmann,H.E. (1942):
Foraminiferal genus Hantkenina and its subgenera . American Journal of Sciences Vol. 240 p. 809-823

Bolli,H.M.; Loeblich,A.R. and Tappan,H. (1957):
The Planktonic foraminiferal families Hantkeninidae, Orbulinidae, Globorotaliidae, and Globotruncanidae . U.S. National Museum Bull. Vol. 215 p. 3-50

Ramsay,W.R. (1962):
Hantkenininae in the Tertiary rocks of Tanganyika . Contributions from the Cushman Foundation for Foraminiferal Research Vol. 13 p. 79-89

Toumarkine,M. (1981):
Discussion de la validité de l'espèce Hantkenina aragonensis Nuttall, 1930. Description de Hantkenina nuttalli n.sp. . Cahiers de Micropaleontologie Vol. 4 p. 109-19

Toumarkine,M. and Luterbacher,H.P. (1985):
Paleocene and Eocene Planktic Foraminifera.
In: Plankton Stratigraphy p. 87-154

Berggren,W.A.; Kent,D.V.; Swisher,C.C. and Aubry,M.P. (1995):
A revised Cenozoic Geochronology and Chronostratigraphy.
In: Geochronology Time Scales and Global Stratigraphic Correlation, SEPM Special Publication Vol. 54

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