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Species Beella digitata Brady 1879


Diagnosis / Definition:
Kennett & Srinivasan (1983):
Test medium to high trochospiral, four to five rapidly increasing chambers in the final whorl, chambers initially spherical, soon becoming ovate and finally radially elon- gate; sutures distinct, depressed; surface consisting of a fine, irregularly cancellate pattern of raised ridges surrounding the circular to subcircular pores; the discrete tubercles surrounding the pores represent spine bases . Aperture interiomarginal, umbilical-extraumbilical, a wide open arch, bordered by a thick lip.
Discussion / Comments:
Kennett & Srinivasan (1983):
The height of the spire in B. digitata shows considerable variation with growth of the individual. There is a tendency toward streptospiral coiling in adult test, suggesting an affinity with Hastigerinella. B. digitata developed from B. praedigitata, from which it differs in having the radially elongate chambers.
Bolli & Saunders (1985):
Banner 2-4; Blow (1960b) described Beella as a subgenus of Globorotalia. Blow (1965) treated it as a subgenus of Globigerina; here it is given full generic rank. Characteristic of the species is the highly trochospiral arrangement, in particular of the chambers forming the early whorls, and the gradually more pronounced elongation of the chambers in the last whorl, with the last chamber becoming quite distinctly pointed. Fully grown specimens with strongly elongate chambers in the last whorl are comparatively rare. More frequent are smaller specimens consisting of the highly trochospiral earlier portion and with the last chambers becoming only slightly elongate. Such forms were included by Parker (1967) in her Globigerina praedigitata. In digitata cf. digitata Blow (1969) placed specimens 'which show the distinct radial elongation of the last two chambers, but which do not have the distinct pointed conical termination' (as in the neotype specimen of digitata). According to Banner 2-4; Blow the position of the aperture is'interiomarginal, umbilical extraumbilical, ventral throughout ontogeny'. In most specimens examined by us the position of the aperture appears virtually umbilical, with only occasionally a trend towards a more peripheral, extraumbilical position. In rare cases, such as in Brady (1884, pt. 8, fig. 8) the aperture even extends beyond the periphery onto the spiral side. Parker (1967) observed praedigitata, first in Zone N 17 (Late Miocene, Globorotalia humerosa Zone), with the position of its real first appearance left uncertain. Blow (1969) quotes its range from either latest N 16 or earliest N 17 to within N 21 (Middle Late Pliocene), and for his cf. digitata from within middle part of N 21 (Late Pliocene) to within N 22 (within Early Pleistocene), whereas he quotes the subspecies digitata originally described from the Holocene - as ranging from N 22 to N 23, i.e. throughout the Pleistocene to Holocene. Observations on DSDP material from Caribbean sites indicate that praedigitata and cf. digitata types continue to be present as immature growth stages of digitata throughout the Pleistocene and Holocene.
Systematics:

1
 Superregnum Eukaryota
  Regnum Protoctista
   Phylum Ciliophora
    Subphylum Postciliodesmatophora
     Ordo Globigerinida
      Superfamilia Globigerinaceae
       Superfamilia Nonionacea
        Familia Globigerinidae
         Genus Beella
          Species Beella digitata

3
  Classis Foraminifera
   Species Beella digitata
    Species Globigerina (Globigerina) quinqueloba
Synonym list:
Kennett & Srinivasan (1983):
1879 Globigerina digitata Brady. - Brady : p.286
p 1884 Globigerina digitata Brady. - Brady : p.599 pl 80 figs. 6-10 (not pl 82 figs 6, 7)
Chaisson & d'Hondt (2000):
1879 Globigerina digitata Brady. - Brady : p.599 pl 80 figs 6-10
1983 Beella digitata Brady. - Kennett & Srinivasan : p.232 pl 58 figs 2, 6-8
Bolli & Saunders (1985):
1884 Globigerina digitata Brady. - Brady : p.599 pl.80, fig.10
1960 Globorotalia (Beela) digitata Bady. - Banner & Blow : p.26 text-figs. 11a-c (=specimen of Brady, 1884, fig. 10, re-illustrated)
1985 Beella digitata Brady. - Bolli & Saunders : p.254 figs. 44.1-4; 7,10,11
Chaisson & Pearson (1997):
1879 Globigerina digitata Brady. - Brady : p.559 pl. 80; fig. 6-10
1983 Beella digitata Brady. - Kennett & Srinivasan : p.232 pl. 58; fig. 2, 6-8
Vincent & Toumarkine (1990):
1879 Globigerina digitata Brady. - Brady : p. 599 pl. 80, figs. 6-10
1990 Beella digitata Brady. - Vincent & Toumarkine : p. 830 pl. 1, fig. 1
Chaisson & Leckie (1993):
1879 Globigerina digitata Brady. - Brady : p.559 pl. 80; fig. 6-10
1983 Beella digitata Brady. - Kennett & Srinivasan : p.232 pl. 58; fig. 2 and 6-8
References:

Brady,H.B. (1879):
Notes on some of the reticularean Rhizopoda of the Challenger Expedition, Part II. Additions to the knowledge of the porcellanous and hyaline types. . Q. J. Microsc. Sci. Vol. 19

Brady,H.B. (1884):
Report on the foraminifera dredged by H.M.S. Challenger during the years 1873-1876.
In: Report of the scientific results of the voyage of H.M.S. Challenger, 1873-1876, Zoology Vol. 9 p. 1-814

Banner,F.T. and Blow,W.H. (1960):
The taxonomy, morphology and affinities of the genera included in the subfamily Hastigerininae . Micropaleontology Vol. 6 p. 19-31

Kennett,J. and Srinivasan,M.S. (1983):
Neogene Planktonic Foraminifera - A Phylogenetic Atlas.

Bolli,H.M. and Saunders,J.B. (1985):
Oligocene to Holocene low latitude planktic foraminifers.
In: Plankton Stratigraphy Eds: Bolli, H.M.Saunders, J.B. p. 155-262

Vincent,E. and Toumarkine,M. (1990):
Neogene planktonic foraminifers from the Western Tropical Indian Ocean, Leg 115.
In: Proceedings of the Ocean Drilling Program, Scientific Results Vol. 115 Eds: Duncan, R.A..Backman, J.Peterson, L.C. p. 795-836

Chaisson,W.P. and Leckie,R.M. (1993):
High-Resolution Planktonic Foraminifer Biostratigraphy of Site 806, Ontong Java Plateau (Western Equatorial Pacific).
In: Proceedings of the Ocean Drilling Program, Scientific Results Vol. 130 Eds: Berger, W.H.Kroenke, L.W..Mayer, L.A..et al. p. 137-178

Chaisson,W.P. and Pearson,P.N. (1997):
Planktonic foraminifer biostratigraphy at Site 925: middle Miocene–Pleistocene.
In: Proceedings of the Ocean Drilling Program, Scientific Results Vol. 154 Eds: Shackleton, N.J.Curry, W.B.Richter, C.Bralower, T.J.

Chaisson,W.P. and d'Hondt,S.L. (2000):
Neogene planktonic foraminifer biostratigraphy at Site 999, Western Caribbean Sea.
In: Proceedings of the Ocean Drilling Program, Scientific Results Vol. 165 Eds: Leckie, R.M.Sigurdsson, H.Acton, G.D.Draper, G. p. 19-56
Anonymous: Unedited TaxonConcept data
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