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Species Voluta triplicata Sowerby 1846

Discussion / Comments:
Griffin & Nielsen (2008):
TYPE MATERIAL. There are two syntypes: NHM-G26412 and NHM-G26413. REMARKS. The figured syntype NHM-G26412 (Sowerby 1846, pl. 4, fig. 74) has three narrow, sharp columellar plaits; many fine, even spiral threads all over; a weak keel at the periphery; quite prominent axial folds all over, commencing halfway down the base; protoconch with a “caricelloid” spine. The other syntype (NHM-G26413) has weaker spiral threads and more prominent axials, a more deeply concave sutural ramp than the figured one and lacks a keel at the periphery. The two specimens may not be conspecific. Del Río & Martínez (2006) figured the syntype NHM-G46412, calling it holotype in the figure captions. This, however, does not serve as a valid lectotype designation. The Chilean species of the genus Proscaphella Ihering, 1907 have been revised by Stuardo & Villarroel (1975). The type species of Proscaphella, Voluta gracilis Philippi, 1887 (i.e., Voluta gracilior Ihering, 1896), belongs in Miomelon Dall, 1907 [type species Volutilithes philippiana Dall, 1890], which makes Proscaphella a synonym of Miomelon. However, contrary to del Río & Martínez (2006), Voluta triplicata does not belong there and the new genus Palaeomelon was introduced by Nielsen & Frassinetti (2007) to accommodate this and related Chilean Neogene volutes. After Nielsen & Frassinetti (2007) went to the printer we discovered homonymy with Voluta triplicata Donovan, 1802, a synonym of Tralia ovula (Bruguière, 1789) (see de Frias Martins 1996), and we propose Palaeomelon jeffi as replacement name for Voluta triplicata Sowerby, 1846. This name is in honour of our friend and colleague Jeffrey (Jeff) Stilwell who contributed much to our knowledge of Southern Hemisphere fossil molluscs.
Synonym list:
Griffin & Nielsen (2008):
1846 Voluta triplicata Sowerby. - Sowerby : p.262 pl. 4; fig. 74 (junior primary homonym of V. triplicata Donovan, 1802)
1887 Voluta triplicata Sowerby. - Philippi : p.66 pl. 7; fig. 8-10
1907 Adelomelon triplicata Sowerby. - Dall : p.365
1907 Proscaphella triplicata Sowerby. - Ihering : p.206
1975 Proscaphella triplicata Sowerby. - Stuardo & Villarroel : 152-153 fig. 19
2006 Miomelon triplicata Sowerby. - del Rio & Martinez : fig. 14.3, 14.4
2007 Palaeomelon triplicata Sowerby. - Nielsen & Frassinetti : p.86 fig. 5.1-5.8
2008 Palaeomelon nom. nov jeffi . - Griffin & Nielsen : pl. 24; fig. 5-8
Was used in synonym list of:
Palaeomelon triplicata Sowerby 1846

Sowerby,G.B.. (1846):
Descriptions of Tertiary fossil shells from South America..
In: Geological observations on South America. Eds: Darwin, C.. p. 249-264

Philippi,R.A.. (1887):
Die tertiären und quartären Versteinerungen Chiles.. p. 266

Ihering,H.. (1907):
Les mollusques fossiles du Tertiaire et du Crétacé Supérieur de l’Argentine . Anales del Museo Nacional de Buenos Aires 3/7 p. 611

Dall,W.H.. (1907):
A review of the American Volutidae . Smithsonian Miscellaneous Collection Vol. 3 p. 341-373

Stuardo,J.. and Villarroel,M.. (1975):
On some living and fossil volutes referred to Miomelon Dall, 1907 and Proscaphella von Ihering, 1907 . The Veliger Vol. 17 p. 139-155

del Rio,C.J.. and Martinez,S.. (2006):
The family Volutidae (Mollusca-Gastropoda) in the Tertiary of Patagonia (Argentina) . Journal of Paleontology Vol. 80 p. 919-945

Nielsen,S.N.. and Frassinetti,D.. (2007):
The Neogene Volutidae (Gastropoda, Neogastropoda) from the Pacific coast of Chile . Journal of Paleontology Vol. 81 p. 82-102

Griffin,M.. and Nielsen,S.N.. (2008):
A revision of the type specimens of Tertiary molluscs from Chile and Argentina described by d’Orbigny (1842), Sowerby (1846), and Hupé (1854) . Journal of Systematic Palaeontology

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