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Species Acarinina angulosa Bolli 1957



Diagnosis / Definition:
Pearson et al. (2006):
DESCRIPTION. Type of wall: Muricate, normal perforate, nonspinose. Test morphology: Low trochospiral test with lobulate peripheral outline; about 10- 1 2 chambers in 2 1/2 whorls; 4 1/2-5 subangular chambers in last whorl gradually increasing in size, separated by straight, radial and depressed sutures; spiral side flat to slightly convex, sutures curved/oblique; umbilicus moderately wide, open, deep; aperture a low interiomarginal-umbilical arch extending nearly to the periphery; axial / edge view subangular. Size: Maximum diameter of holotype 0.40 mm, thickness 0.28 mm.
Discussion / Comments:
Van Eijden & Smit (1991):
Remarks. See A. coalingensis (Cushman and Hanna, 1927).
Pearson et al. (2006):
DISTINGUISHING FEATURES.- The diagnostic features of Acarinina angulosa are the strongly angular disposition of the chambers of the last whorl and the strongly lobulate peripheral outline. DISCUSSION.- This distinctive taxon is a common component of lower Eocene (sub)tropical assemblages where it is associated with other acarininids (soldadoensis, interposita, pentacamerata, pseudotopilensis) and morozovellids (suhbotinae, formosa, gracilis, aragonensis). In the course of our studies in Egypt we have found that this taxon has its FAD just below the base of the CIE / PETM interval, i.e., within the uppermost part of Zone P5 (as redefined in Berggren and Pearson, 2005). PHYLOGENETIC RELATIONSHIPS.- This species evolved from Acarinina soldadoensis in uppermost Zone P5 and probably gave rise to A. cuneicamerata. STRATIGRAPHIC RANGE.- Zone P5 (uppermost part) to Zone E7. Note that the Paleocene part of the range was not recorded by Olsson and others (l 999). GEOGRAPHIC DISTRIBUTION.- Ranges from tropical to temperate regions; Arabian Sea, north Pacific Ocean (DSDP Site 47), New Jersey coastal Plain (Bass River), Tethyan deposits of northern Africa (Egypt) and occurrences in Spain (Alicante Province) and New Guinea. STABLE ISOTOPE PALEOBIOL0GY.- No data available.
Systematics:

32
 Familia Globorotaliidae
  Genus Acarinina
   Species Acarinina angulosa

35
  Ordo Foraminiferida
   Superfamilia Globigerinaceae
    Familia Truncorotaloididae
     Genus Acarinina
      Species Acarinina angulosa
Synonym list:
Van Eijden & Smit (1991):
1957 Globigerina soldadoensis angulosa Bolli. - Bolli : p.71 pl. 16, figs. 4-6
1991 Acarinina angulosa Bolli. - Van Eijden & Smit : p.111
Pearson et al. (2006):
1957 Globigerina soldadoensis angulosa Bolli. - Bolli : p.71 pl. 16; fig. 4-6 [lower Eocene Globorotalia formosa formosa Zone, Lizard Springs area, Trinidad]
1957 Globigerina soldadoensis angulosa Bolli. - Bolli : p.162 pl. 35; fig. 8a-c [lower Eocene Globorotalia palmerae Zone, Navet Fm., Trinidad]
1965 Acarinina soldadoensis angulosa Bolli. - Hillebrandt : p.345 pl. 5; fig. 11 [Zone P9, Agost, Alicante Province, Spain]
? 1972 Turborotalia (Acarinina) soldadoensis angulosa Bolli. - Samuel et al. : p.190 pl. 68; fig. 1a-c [lower Eocene Globorotalia subbotinae Zone, Hradisko, Middele Vah Valley, West Carpathian]
1974 Acarinina soldadoensis angulosa Bolli. - Fleisher : p.1014 pl. 4; fig. 1 [lower Eocene Zone P9, DSDP Site 220, Arabian Sea]
1979 Muricoglobigerina soldadoensis angulosa Bolli. - Blow : p.1122 pl. 109, fig. 9; pl. 131, fig. 4-5 [lower Eocene Zone P6 of Blow, 1979), DSDP Hole 47.2, Shatsky Rise, north-west Pacific Ocean]
? 1984 Muricoglobigerina soldadoensis angulosa Bolli. - Belford : p.30 pl. 22; fig. 9-12 [lower Eocene Zones PS-9, Papua New Guinea]
2006 Acarinina angulosa Bolli. - Pearson et al. : p.268 pl. 9.3; fig. 11-16 (Pl. 9.3, Figs. 13-15: new SEMs of holotype of Globigerina soldadoensis angulosa Bolli)
Specimen:
Smithsonian Museum of Natural History, Washington, D.C., Inventory number: USNM P5037
References:

Bolli,H.M. (1957):
The genera Globigerina and Globorotalia in the Paleocene-Lower Eocene Lizard Springs Formation of Trinidad, B.W.I . Bulletin of the U.S. National Museum Vol. 215 p. 61-82

Bolli,H.M. (1957):
Planktonic Foraminifera from the Eocene Navet and San Fernando formations of Trinidad, B.W.I. . Bull. U.S. natl. Mus. Vol. 215 p. 155-172

Hillebrandt,v.A.. (1965):
Los foraminiferos planctonicos, nummulitidos y coccolitoforidos de la zona de Globorotalia palmerae del Cuisiense (Eoceno Inferior) en el SE de Espania, (provincias de Murcia y Alicante) . Revista Espanola de Micropaleontologia Vol. 8 p. 323-394

Samuel,O..; Borza,K.. and Kohler,E.. (1972):
Microfauna and lithostratigraphy of the Paleogene and adjacent Cretaceous of the Middle Vah Valley (West Carpathian) . Geologicky ustav Dionyza Stura, Bratislava p. 246

Fleisher,R.L. (1974):
Cenozoic planktonic foraminifera and biostratigraphy, Arabian Sea, Deep Sea Drilling Project, Leg 23A.
In: Initial Reports of the Deep Sea Drilling Project Vol. 23 Eds: Whitmarsh, R.B.Weser, O.E.Ross, D.A. p. 1001-1072

Blow,W.H. (1979):
The Cainozoic Globigerinida. 3 Vols p. 1413 pp

Belford,D.J. (1984):
Tertiary Foraminifera and Age of Sediments, Ok Tedi-Wabag, Papua, New Guinea . BureauMineral Resources, Geology and Geophysics, Australia Bulletin Vol. 216

Van Eijden,A.J.M. and Smit,J. (1991):
Eastern Indian Ocean Cretaceous and Paleogene quantitative biostratigraphy.
In: Proceedings of the Ocean Drilling Program, Scientific Results Vol. 121 Eds: Weissel, J.Peirce, J.Taylor, E.Alt, J. p. 77-123

Pearson,P.N.; Olsson,R.K.; Hemleben,C.; Huber,B.T. and Berggren,W.A. (2006):
Atlas of Eocene Planktonic Foraminifera. p. 1-513

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