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Species Haeuslerina helvetojurassica Haeusler 1881 | ||||||
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| Discussion / Comments: | ||||||
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Boudagher-Fadel et al. (1997): Remarks: Haeusler (1881b) stated that the type level for his new species was the zone of 'Ammonites transversarius' in the Birmensdorfer Schichten of the Jurassic, near Buren (Canton Aargau), Switzerland. In his doctoral dissertation (1881a), however, no type locality was designated. Oesterle (1968) designated the 'Eisengraben' section (80 km NW of Zurich, Canton Aargau) of the Birmensdorfer Schichten as the type locality for all foraminifera species described by Haeusler from those beds. The topotypes illustrated here (Plate 2.6, Figs 1-8) were collected by Stam (1986, p. 111) from the Eisengraben section, the same layer as Oesterle's sample no. 4; 'they are rare and small in size, [and] this sample contained well preserved planktonics'. Earlier, Oesterle (1969) had attempted to revise the work of Haeusler and redescribed 'G.' helvetojurassica from the Renggeri Tonen of Liesberg (75 km west of the Eisengraben section). These were clearly pyrite moulds as were Masters' (1977, plate 21, figs 2-4) illustrated SEM images (called 'Globigerina? helvetojurassica') from the Oxfordian of the Birmensdorfer Beds of Eisengraben. In contrast, the specimens from the Eisengraben section collected by Stam (1986, plate 7, figs 6-12; plate 8, figs 1-3) from the Birmensdorfer Schichten, Transversarium Zone ('Middle' Oxfordian) were perfectly preserved. Some of Stam's figured specimens have been re-imaged here (Plate 2.6, figs 1-8). All are covered in scattered pseudomuricae, not aligned into ridges, and very distinctively possess very small umbilici and primary apertures which were clearly intraextraumbilical. Stam (1986, pp. 110, 111) did not notice the difference in the extent of the aperture in his specimens compared to that of Globuligerina oxfordiana (Grigelis), and he mistakenly considered the topotypes of 'G.' helvetojurassica to be synonymous with G. oxfordiana. As Globigerina helvetojurassica Haeusler was published in 1881, while G. oxfordiana Grigelis was not published until 1958, the former name would have been the senior one; Stam endeavoured to overcome this by referring Haeusler's name to the International Commission of Zoological Nomenclature, claiming that G. helvetojurassica should be regarded as nomen oblitum (a forgotten name, ICZN article 23b) and not used. However, the case was not followed up by Stam, and the ICZN did not accept it (Dr P. Tubbs (ICZN), personal communication). Now we know that G. helvetojurassica Haeusler belongs to a genus (Haeuslerina gen.nov.) distinct from Globuligerina. The specimens figured by Masters (1977, pI. 21, figs 2-4) and Oesterle (1968, text-figs 51, 52a) were internal moulds, probably like the specimens in Haeusler's own collection (Masters, 1977, p. 459). Therefore, the specimens figured for this chapter, like those figured by Stam (1986, plate 7, figs 6-12, plate 8, figs 1-3) are those which must now be used for the interpretation of the morphology of Haeuslerina helvetojurassica (Haeusler). Although Haeusler (188la; l881b) did not choose a holotype for his species, the existence of syntypes (referred to by Masters, 1977), albeit internal moulds, unfortunately makes it impossible to designate a neotype (ICZN, article 75). |
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| Systematics: | ||||||
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22 Subordo Globigerinina Superfamilia Favusellacea Familia Conoglobigerinidae Genus Haeuslerina Species Haeuslerina helvetojurassica |
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| Synonym list: | ||||||
| Boudagher-Fadel et al. (1997): 1881 Globigerina helvetojurassica Haeusler. - Haeusler : p.36 pl. 2; fig. 44, 44a
p 1986 Globuligerina oxfordiana Grigelis. - Stam : pl. 7, fig. 6-12 (not fig. 1-5);
pl. 8, fig. 1-3 (not pl. 9, fig. 1-5;
pl. 14, fig. 3, 4, 8-15)
1997 Haeuslerina helvetojurassica Haeusler. - Boudagher-Fadel et al. : p.43 pl. 2.6; fig. 1-8; Figure: 2.2
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| References: | ||||||
Haeusler,R.. (1881): Stam,B.. (1986): Boudagher-Fadel,M.K.; Banner,F.T.; Whittaker,J.E. and Simmons,M.D. (1997): |
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| Anonymous: Unedited TaxonConcept data | ||||||
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