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Species Subbotina hornibrooki Brönniman 1952


Diagnosis / Definition:
Pearson et al. (2006):
DESCRIPTION. Type of wall: Coarsely cancellate, sacculifer-type wall texture, normal perforate, spinose. Test morphology: Test low trochospiral, globular, slightly lobulate in outline, chambers globular, embracing; in spiral view 3 1/2 to nearly 4 globular, embracing chambers in ultimate whorl, increasing rapidly in size, sutures slightly depressed, straight, ultimate chamber reduced in size and directed over the umbilicus; in umbilical view 31h to nearly 4 globular, embracing chambers, increasing rapidly in size, sutures slightly depressed, straight, umbilicus a small opening, enclosed by surrounding chambers and partly closed by the projection of the ultimate chamber, aperture umbilical, bordered by a narrow continuous, lip, ultimate chamber reduced in size relative to the penultimate chamber, in edge view chambers globular in shape, embracing. Size: Maximum diameter of holotype 0.25 mm, thickness 0. 19 mm.
Discussion / Comments:
Pearson et al. (2006):
DISTINGUISHING FEATURES.- Subbotina hornibrooki is distinguished by its compact, low trochospiral test with 3 1/2 to nearly 4 globular embracing chambers in the ultimate whorl, reduced ultimate chamber that is directed over the umbilicus, and its coarsely cancellate sacculifer-type wall texture. DISCUSSION.- Subbotina hornibrooki has been misinterpreted by workers probably due to the line drawing of the holotype, which presents a generalized morphology. Brönnimann reported the wall to be finely perforate, which somewhat confuses matters. The holotype illustrated for the first time in SEM (Pl. 6.12, Figs. 1-3) shows a coarsely perforate sacculfer-type wall texture that places this species in the Subbotina velascoensis group of species. Most of the specimens illustrated by Blow (1979) lack this diagnostic wall texture. Blow noted the close affinity of Globigerina finlayi Brönnimann, which was described in the same paper as hornibrooki, and we here consider it a synonym. The holotype of Globigerina bacuana Khalilov has been viewed by WAB and questionably referred to this species. PHYLOGENETIC RELATIONSHIPS.- Subbotina hornibrooki belongs to the Subbotina velascoensis group of species that developed in the Paleocene from Subbotina cancellata Blow. Subbotina hornibrooki developed from Subbotina velascoensis in Zone P5 by developing a more compact test with a reduced ultimate chamber that is directed over the umbilicus. GEOGRAPHIC DISTRIBUTION.- Global in low to mid latitudes. STABLE ISOTOPIC PALEOBIOL0GY.- No data available.
Systematics:

35
 Ordo Foraminiferida
  Superfamilia Globigerinaceae
   Familia Globigerinidae
    Genus Subbotina
     Species Subbotina hornibrooki
Synonym list:
Pearson et al. (2006):
1952 Globigerina hornibrooki Brönnimann. - Brönnimann : p.15 pl. 12; fig. 4-6 (holotype) [upper Paleocene Zone P5, Lizard Springs Fm., Trinidad]
1952 Globigerina finlayi Brönnimann. - Brönnimann : p.18 pl. 2; fig. 10a-c [middle Eocene Lizard Springs Fm., Trinidad]
? 1956 Globigerina bacuana Khalilov. - Khalilov : p.235 pl. 3; fig. 4a-c [upper Paleocene, Maly Caucasus, Azerbaidzhan]
1979 Subbotina hornibrooki Brönniman. - Blow : p.1269 pl. 124; fig. 8 [lower Eocene Zone E5, DSDP Hole 47.2, northwest Pacific Ocean]
p 1979 Subbotina hornibrooki hornibrooki Brönnimann. - Blow : p.1269 pl. 101; fig. 7 [upper Paleocene Zone P5, DSDP Hole 47.2, northwest Pacific Ocean]. [Not pl. 124: fig. 7; p1.142: fig. 6; pl. 160: fig. 4.]
non 1995 Subbotina hornibrooki Brönniman. - Poag & Commeau : p.144 pl. 3; fig. 25, 26 [= Subbotina roesnaesensis n. sp.]
2006 Subbotina hornibrooki Brönniman. - Pearson et al. : p.140 pl. 6.12; fig. 1-16 (Pl. 6.1 2, Figs. 1-3: new SEMs of holotype of Globigerina hornibrooki Bronnimann)
Specimen:
Smithsonian Museum of Natural History, Washington, D.C., Inventory number: USNM 370078
References:

Brönnimann,P. (1952):
Trinidad Paleocene and lower Eocene Globigerinidae . Bulletins of American paleontology Vol. 34(143) p. 1-34

Khalilov,D.M. (1956):
O pelagicheskoy faune foraminifer Paleogenovykh otlozheniy Azerbaydzhana . Trudy Instituta Geologii, Akademiya Nauk Azerbaydzhanskoy SSR Vol. 17

Blow,W.H. (1979):
The Cainozoic Globigerinida. 3 Vols p. 1413 pp

Poag,C.W.. and Commeau,J.A.. (1995):
Paleocene to middle Miocene planktic foraminifera of the southwestern Salisbury Embayment, Virginia and Maryland: biostratigraphy, allostratigraphy, and sequence stratigraphy . Journal of Foraminiferal Research Vol. 25 p. 134-155

Pearson,P.N.; Olsson,R.K.; Hemleben,C.; Huber,B.T. and Berggren,W.A. (2006):
Atlas of Eocene Planktonic Foraminifera. p. 1-513
Anonymous: Unedited TaxonConcept data
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