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Species Acarinina pseudosubsphaerica Pearson & Berggren 2006


Diagnosis / Definition:
Pearson et al. (2006):
DESCRIPTION. Type of wall: Weakly muricate, cancellate on spiral side with large, bladed muricae on umbilical side, normal perforate, nonspinose. Test morphology: Compact subspherical test coiled in a high trochospire of about two whorls with circular, slightly lobate outline; 4-6 moderately inflated, radially compressed chambers in final whorl; dorsal sutures wide and depressed, between circular chambers; primary aperture slit-like or a low, broad arch, situated in a wide, open umbilicus, and often covered by a small, sparsely muricate final chamber. Size: Maximum diameter of holotype 0.19 mm, thickness 0.16 mm.
Discussion / Comments:
Pearson et al. (2006):
ETYMOLOGY.- Named for its resemblance to the Paleocene species Acarinina subsphaerica Subbotina. DISTINGUISHING FEATURES.- Acarinina pseudosubsphaerica n. sp. is distinguished from all other middle Eocene acarininids by its high spire, small size and subspherical shape. It is similar in morphology to the Paleocene species, A. subsphaerica (Subbotina), but differs from that form and A. alticonica Fleisher by having less closely appressed chambers and a broader umbilicus. DISCUSSION.- Despite being a relatively common form in the lower and middle Eocene, Acarinina pseudosubsphaerica n. sp. has not been formally recorded. Acarinina alticonica Fleisher is closely related and the two species have been observed to intergrade in Zone E6/E7 (undifferentiated) of Tanzania Drilling Project Site 2 (P.N. Pearson, unpublished data). Lu and Keller (1995) recorded a first occurrence in Zone P9 (=E7) of Acarinina aquiensis in DSDP Site 577 (Shatsky Rise, north-west Pacific Ocean), which is regarded as conspecific here, and not referable to Loeblich and Tappan's species (which is a Paleocene form from the Aquia Formation of Virginia and Maryland probably related to/conspecific with Acarinina subsphaerica; see Berggren and Norris, 1 997). PHYLOGENETIC RELATIONSHIPS.- Descended from A. alticonica. STRATIGRAPHIC DISTRIBUTION.- Zone E7 to Zone E10 GEOGRAPHIC DISTRIBUTION.- Probably widely distributed in tropical and mid-latitude sites. STABLE ISOTOPE PALEOBIOL0GY.- Recorded (under the name Muricoglobigerina esnehensis) as a mixed layer dwelling species by Pearson and others, (1993).
Systematics:

35
 Ordo Foraminiferida
  Superfamilia Globigerinaceae
   Familia Truncorotaloididae
    Genus Acarinina
     Species Acarinina pseudosubsphaerica
Synonym list:
Pearson et al. (2006):
1993 Muricoglobigerina esnehensis Nakkady. - Pearson et al. : pl. 1; fig. 20 [middle Eocene Zone P 11 -P12, DSDP Site 523, Walvis Ridge, South Atlantic Ocean]; [Not Nakkady, 1950]
1995 Acarinina aquiensis Loeblich & Tappan. - Lu & Keller : pl. 2; fig. 24-26 [Lower Eocene Zone P9, DSDP Site 577, Shatsky Rise, north-west Pacific Ocean]; [Not Loeblich and Tappan, 1957]
2004 Acarinina cf. subsphaerica Subbotina. - Pearson et al. : p.37 pl. 2; fig. 4 [middle Eocene, Zone P1 l , Tanzania Drilling Project Site 2, Kilwa Masoko, Tanzania]; [Not Subbotina, 1947]
2006 Acarinina n sp. pseudosubsphaerica Pearson & Berggren. - Pearson et al. : p.267 pl. 9.2; fig. 9-16 (Pl. 9.2, Figs. 9- 10, 14- 15: SEMs of holotype and paratypes)
Specimen:
Smithsonian Museum of Natural History, Washington, D.C., Inventory number: USNM 523427
Smithsonian Museum of Natural History, Washington, D.C., Inventory number: USNM 523427a
Smithsonian Museum of Natural History, Washington, D.C., Inventory number: USNM 523427b
Smithsonian Museum of Natural History, Washington, D.C., Inventory number: USNM 523427c
References:

Pearson,P.N.; Shackleton,N.J. and Hall,M.A. (1993):
The Stable Isotope Paleoecology of Middle Eocene Planktonic Foraminifera and Multi-species integrated Isotope Stratigraphy . Journal of Foraminiferal Research Vol. 23 p. 123-140

Lu,G. and Keller,G. (1995):
Planktic Foraminiferal Faunal Turnovers in the Subtropical Pacific during the Late Paleocene to Early Eocene . Journal of Foraminiferal Research Vol. 25 p. 97-116

Pearson,P.N.; Nicholas,C.J..; Singano,J.M..; Bown,P.R..; Coxali,H.K..; van Dongen,B.E..; Huber,B.T.; Karega,A..; Lees,J.A..; Misaky,E..; Pancost,R.D..; Pearson,M.. and Roberts,A.P.. (2004):
Paleogene and Cretaceous sediment cores from the Kilwa and Lindi areas of coastal Tanzania: Tanzania Drilling Project Sites 1-5 . Journal of African Earth Sciences Vol. 39 p. 25-62

Pearson,P.N.; Olsson,R.K.; Hemleben,C.; Huber,B.T. and Berggren,W.A. (2006):
Atlas of Eocene Planktonic Foraminifera. p. 1-513
Anonymous: Unedited TaxonConcept data
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