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Species Acarinina quetra Bolli 1957


Diagnosis / Definition:
Pearson et al. (2006):
DESCRIPTION. Type of wall: Moderately to strongly muricate with concentration of muricae along peripheral margin of last whorl, nonspinose, normal perforate. Test morphology: Low trochospiral; test subquadrate, planoconvex; peripheral margin lobulate; chambers on umbilical side moderately inflated, flattened on spiral side; 4 broadly subtriangular chambers visible in umbilical view; anterior and posterior margins of last chamber flattened resulting in disjunct geometry with bordering chambers and conical apex of last chamber; sutures depressed/incised, curved to sinuous, radial; umbilicus relatively wide, deep; aperture an umbilical-extraumbilical, arch extending towards (but not to) the peripheral margin; 10-12 chambers in 2 1/2-3 whorls on spiral side; chambers loosely disposed, meeting almost at right angles, increasing gradually in size; final chamber sometimes reduced in size; chambers lens-shaped, tangentially longer than radially broad in general, overlapping; sutures curved; discrete intercameral openings visible on some individuals (depending on preservation); planoconvex in edge view, final chamber distinctly anguloconical with high conical angle; periphery distinctly, but discontinuously, muricocarinate Size: Maximum diameter of holotype 0.50 mm, thickness 0.38 mm.
Discussion / Comments:
Pearson et al. (2006):
DISTINGUISHING FEATURES.- This taxon is distinguished by its angular test whose chambers are loosely disposed sequentially to each other at - 90° and with a distinct, but discontinuous, peripheral muricocarina. DISCUSSION.- Acarinina quetra is a distinctive form that with its cuneate chambers and peripheral concentration of muricae, bears a superficial resemblance (in some instances) to the stratigraphically younger (middle Eocene) A. topilensis. PHYLOGENETIC RELATIONSHIPS.- Descended from A. pseudotopilensis. STRATIGRAPHIC RANGE.- Zone E3 (upper part) to Zone E6. GEOGRAPHIC DISTRIBUTION.- Widely distributed in (sub)tropical regions (Caribbean, Atlantic, Indo- Pacific, Tethyan/Mediterranean regions, North Caucasus) STABLE ISOTOPE PALEOBIOLOGY.- No data available.
Systematics:

35
 Ordo Foraminiferida
  Superfamilia Globigerinaceae
   Familia Truncorotaloididae
    Genus Acarinina
     Species Acarinina quetra
Synonym list:
Pearson et al. (2006):
1957 Globorotalia quetra Bolli. - Bolli : p.79 pl. 19; fig. 1-3 (holotype), 4-6 (paratype) [Globorotalia formosa formosa Zone, upper Lizard Springs Fm., Trinidad]
1962 Globorotalia (Acarinina) quetra Bolli. - Hillebrandt : p.144 pl. 14; fig. 2a-c [Zone G, Reichenhall Basin, south-west Germany]
1974 Acarinina quetra Bolli. - Fleisher : pl. 3; fig. 3 [Zone P8, DSDP Site 220, Arabian Sea]
1975 Globorotalia quetra Bolli. - Stainforth et al. : p.221 text-fig. 80.1-6 [G. formosa Zone, Lizard Springs Fm., Trinidad]; 4-6 (from Bolli, 1957a, reillustrated)
1977 Acarinina quetra Bolli. - Berggren : p.251 chart no. 10 (reillustrations from literature)
1979 Globorotalia (Truncorotaloides) quetra Bolli. - Blow : p. 1034-1036 pl. 122, fig. 4, 7-9; pl. 200, fig. 5, 6; pl. 123, fig. 6-9; pl. 202, fig. 1-6 [lower Eocene Zone P7, DSDP Site 47, Shatsky Rise, north-west Pacific Ocean]; pl. 129, fig. 5 [lower Eocene Zone P8a, DSDP Site 47, Shatsky Rise, northwest Pacific Ocean]; pl. 132, fig. 7-9; pl. 201, figs. 4-6 [lower Eocene Zone P7, DSDP Site 47, Shatsky Rise, northwest Pacific Ocean]; pl. 140: fig. 4 [lower Eocene Zone P8b, DSDP Site 20, South Atlantic Ocean] (note p. 1036 misprinted as 1136)
1985 Morozovella quetra Bolli. - Snyder & Waters : p.448 pl. 9; fig. 4-6 [lower Eocene Zone P6, DSDP Site 550, Porcupine Abyssal Plain, north-east Atlantic Ocean]
2000 Acarinina quetra Bolli. - Warraich et al. : p.193 fig. 17.13-15 [Zone P7, Rakhi Nala section, Sulaiman Range, Pakistan]
2001 Acarinina quetra Bolli. - Warraich & Ogasawara : p.29 fig. 7.4-6 [Zone P7, Rakhi Nala section, Sulaiman Range, Pakistan]
2006 Acarinina quetra Bolli. - Pearson et al. : p.309 pl. 9.19; fig. 1-16 (Pl. 9.19, Figs. 1-3: new SEMs of Globorotalia quetra Bolli)
Specimen:
Smithsonian Museum of Natural History, Washington, D.C., Inventory number: USNM P5070
Smithsonian Museum of Natural History, Washington, D.C., Inventory number: USNM P5071
References:

Bolli,H.M. (1957):
The genera Globigerina and Globorotalia in the Paleocene-Lower Eocene Lizard Springs Formation of Trinidad, B.W.I . Bulletin of the U.S. National Museum Vol. 215 p. 61-82

Hillebrandt,v.A.. (1962):
Das Paleozän und seine Foraminiferenfauna im Becken von Reichenhall und Salzburg . Bayerische Akademie Wissenschaft, Mathematischen-Naturwissenschaften Klasse Vol. 108 p. 188 p.

Fleisher,R.L. (1974):
Cenozoic planktonic foraminifera and biostratigraphy, Arabian Sea, Deep Sea Drilling Project, Leg 23A.
In: Initial Reports of the Deep Sea Drilling Project Vol. 23 Eds: Whitmarsh, R.B.Weser, O.E.Ross, D.A. p. 1001-1072

Stainforth,R.M.; Lamb,J.L.; Luterbacher,H.P.; Beard,J.H. and Jeffords,R.M. (1975):
Cenozoic planktonic foraminiferal zonation and characteristics of index forms . Paleontological ContributionsArticle 62 p. 425

Berggren,W.A. (1977):
Atlas of Paleogene planktonic foraminifera. Some species of the genera Subbotina, Planorotalites, Morozovella, Acarinina and Truncorotaloides.
In: Oceanic Micropaleontology Eds: Ramsay, A.T.S. p. 205-299

Blow,W.H. (1979):
The Cainozoic Globigerinida. 3 Vols p. 1413 pp

Snyder,S.W. and Waters,V.J. (1985):
Cenozoic planktonic foraminiferal biostratigraphy of the Goban Spur region.
In: Deep Sea Drilling Project Vol. 80

Warraich,M.Y..; Ogasawara,K.. and Nishi,H.. (2000):
Late Paleocene to early Eocene planktonic foraminiferal biostratigraphy of the Dungan Formation, Sulaiman Range, Central Pakistan . Paleontological Research Vol. 4 p. 275-301

Warraich,M.Y.. and Ogasawara,K.. (2001):
Tethyan Paleocene-Eocene planktic foraminifera from the Rakhi Nala and Zinda Pir land sections of the Sulaiman Range, Pakistan . Science Reports of the Institute of Geoscience University of Tsukuba, section B Vol. 22 p. 1-59

Pearson,P.N.; Olsson,R.K.; Hemleben,C.; Huber,B.T. and Berggren,W.A. (2006):
Atlas of Eocene Planktonic Foraminifera. p. 1-513
Anonymous: Unedited TaxonConcept data
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