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Species Pseudokossmaticeras duereri Redtenbacher 1873



Diagnosis / Definition:
Ward & Kennedy (1993):
Description.- Holotype is crushed internal mold, distorted into ellipse, with maximum diameter of 71 mm; all of inner whorls missing. Coiling appears to have been serpenticone, with wide umbilicus of moderate depth. Because of crushing, original form of whorl section unclear, although umbilical wall rounded and undercut, flanks broadly rounded and venter rounded. Estimated five deep, strong, narrow, straight and prorsiradiate constrictions per whorl. They are preceded by strong, parallel, rounded rib with distinct, sharp umbilical bulla, and followed by similar, but somewhat weaker rib. On first half of outer whorl, which is rather badly deformed, there are an estimated 18 ribs between two conspicuous constrictions. As preserved, these are prorsiradiate, feebly flexed, and arise in groups of two or three from small umbilical bullae. Occasional long, non-bullate ribs extend to umbilical seam, and there are also shorter, intercalated ribs. On last part of outer whorl, there are 12 ribs between last two constrictions; all long, feebly prorsiradiate, and most bear feeble umbilical bulla. Sutures undecipherable. Small Biscay specimen is in form of silicone squeeze taken from a natural mold, some 43 mm in diameter. Outer whorl corresponds closely to that of holotype, final section showing primary ribs arising singly from feeble umbilical bullae whereas preceding part has ribs arising both singly and in pairs. Inner whorls, not preserved in holotype, show that umbilical bullae, 9-10 per half whorl, were relatively prominent, and ribs weak. Much larger is UWBM 76728, some 65 mm in diameter, with crowded prorsiradiate ribs, great majority primaries, straight on flank, but projected strongly forward on ventrolateral shoulder, with maximum development across venter. Periodic constrictions associated with strong collar-ribs.
Discussion / Comments:
Ward & Kennedy (1993):
#Type. - Holotype is the original of Redtenbacher, 1873, p. 106, Pl. 24, fig. l a d , No. 1873/01/22, in the collections of the Austrian Geological Survey, from the Gosau Beds of Gosau, Austria. Discussion. - Redtenbacher's figure is accurate, if reversed. Pseudokossmaticeras duereri most closely resembles P. pacificum (Stoliczka, 1865, p. 160, PI. 77, fig. 9), a species in which all the ribs are long, with no branching or intercalation, save near constriction, whilst they follow a somewhat concave course. Madagascan examples illustrated by Collignon (1971, Pl. 64, fig. 2398, Pl. 653, fig. 2418) show these features well, and are somewhat sparser ribbed than is P. duereri. Pseudokossmaticeras brandti (Redtenbacher, 1873) (p. 106, Pl. 24, fig. 1; see Thiedig and Wiedmann, 1976) is coarsely and sparsely ribbed at comparable diameters, as in P. aturicus (Seunes, 189 1, p. 17, Pl. 15(6), figs. 2,3), a synonym, which also possesses more regularly bifurcating ribs. In P. galicianum (Favre, 1869) (p. 16, Pl. 3, fig. 5a, b; see also Thiedig and Wiedmann, 1976, p. 17, Pl. 2 figs. 1, 3), of which P. tchihatcheffi (Böhm, 1927) (p. 2 17, Pl. 13, fig. 1) may be a synonym, constrictions are less obvious, and there is a higher whorl section, smaller umbilicus, and obvious alternation of long and short ribs at smaller sizes (e.g., Naidin and Shimanskij, 1959, Pl. 13, fig. 1). Pseudokossmaticeras tercense (Seunes, 189 1) (p. 16, Pl. 15(6), fig. 4; see above, and Thiedig and Wiedmann, 1976, p. 18, Pl. 1, fig. 2, Pl. 2, fig. 2) may be similarly distinguished, and has many more branching ribs. Pseudokossmaticeras muratovi Mikhailov, 195 1 (p. 77, Pl. 13, fig. 52) is also more coarsely ribbed, the ribs branching high on the flank. Pseudokossmaticeras cerevicianum (Petho, 1906) (p. 95, Pl. 6, figs. 2, 3) is much more coarsely ribbed and bullate. Occurrence. - The Biscay specimens are from Member II (lower Maastrichtian) and from just beneath the K-T boundary (Member V, upper Maastrichtian) at Bidart, France. The age of the holotype from Gosau, Austria, is not known.
Systematics:

38
 Ordo Ammonoidea
  Subordo Ammonitina
   Superfamilia Desmocerataceae
    Familia Kossmaticeratidae
     Subfamilia Kossmaticeratinae
      Genus Pseudokossmaticeras
       Species Pseudokossmaticeras duereri
Synonym list:
Ward & Kennedy (1993):
1873 Ammonites düreri Redtenbacher. - Redtenbacher : p.118 pl. 7; fig. 20
1907 Holodiscus düreri Redtenbacher. - Wisniowski : p.196
1925 Kossmaticeras düreri Redtenbacher. - Diener : p.98
1958 Pseudokossmaticeras duereri Redtenbacher. - Reyment : p.33 pl. 7; fig. 1
1993 Pseudokossmaticeras duereri Redtenbacher. - Ward & Kennedy : p. 16, 27 fig. 16.2, 16.3, 25.6
Stratigraphy - relative ages:
upper Maastrichtian - lower Maastrichtian: Ward & Kennedy (1993)
References:

Redtenbacher,A.. (1873):
Die Cephalopodenfauna der Gosauschichten in den nordöstlichen Alpen: . Abhandlungen der königlich-kaiserlichen geologischen Reichsanstalt Vol. 5 p. 91-140

Wisniowski,T.. (1907):
Über die Obersenone flyschfauna von Leszczyny . Beitr. Paläont. Geol. Öst.-Ung. Vol. 30 p. 191-205

Diener,C.. (1925):
Ammonoidea neocretacea . Fossilium Cat, (1: Animalia) Vol. 29 p. 244 pp.

Reyment,R.A. (1958):
Neubeschreibung der Redtenbacher'schen ammoniten- originale aus den Gosauschichten . Stockholm Contributions to Geology Vol. 2 p. 31-49

Ward,P.D.. and Kennedy,W.J.. (1993):
Maastrichtian Ammonites from the Biscay Region (France, Spain) . Memoir (The Paleontological Society), Journal of Paleontology34 (Supplement to Vol. 67)(5) p. 1-58

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