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Species Eubaculites labyrinthicus Morton 1834



Diagnosis / Definition:
Cobban & Kennedy (1995):
DESCRIPTION.- A rare species. The authors have seen only four fragments from the Prairie Bluff Chalk in addition to the types. All are phosphatic internal molds that range from 8-17 mm whorl height. Lectotype (Morton, 1834, plate 13, figure 10) is septate fragment 35.8 mm long. Taper angle 5 degrees; whorl section compressed with intercostal whorl breadth to height ratio 0.58. Dorsum flattened and barely convex. Dorsolateral margin narrowly rounded, dorsal flanks flattened, midflank region broadly rounded, ventral flanks flattened and convergent, venter fastigiate. Strong nodes, elongated parallel to length of shell, perch on dorsalmost flank, two occumng in distance equal to whorl height. Low, broad rib connects each node to a similarly elongated but slightly obliquely aligned midlateral tubercle. Midlateral tubercles give rise to delicate riblets and striae, also present on interspaces, and projected forward to intersect line of venter at acute angle. Dorsum decorated by feebly convex ribs and striae. Suture has broad, bifid, rectangular saddles, L/U with variable degree of incision, E/L narrow and bifid, U broad and bifid.
Discussion / Comments:
Cobban & Kennedy (1995):
TYPES.- Lectotype, here designated, is number 72868 in the collection of the Academy of Natural Sciences of Philadelphia, the original of Morton, 1834, plate 13, figure 10. Paralectotype is no. 72869 in the same collection. Both are from the Prairie Bluff Chalk of Alabama. MATERIAL.- Four specimens, USNM 463170-463173. DISCUSSION.- The most important synonym of E. labyrinthicus is Baculites vagina var. ootacodensis Stoliczka, 1866. The lectotype (designated by Kennedy, 1986d, page 193) is the original of Stoliczka's plate 90, figure 14, a cast of which is before us. It is a wholly septate internal mold, 56 mm long, with a maximum preserved whorl height of 15.5 mm. The intercostal whorl breadth to height ratio is 0.62, the dorsum broad and very slightly convex to flat. The dorsolateral margin is narrowly rounded, the dorsal flanks flattened in intercostal section, the mid-flank region broadly rounded; the ventral flanks are flattened and converge to a bluntly fastigiate venter. Two ribs occur in a distance equal to the whorl height, parts of four ribs being preserved on the specimen. They begin at small dorsolateral nodes, perched on the dorsolateral margin and elongated parallel to the length of the shell. Ribs are coarse, transverse and slightly prorsiradiate on the inner flank before strengthening into a midlateral tubercle that is elongated in a slightly prorsiradiate direction. These tubercles give rise to a progressively weakening prorsiradiate rib that slants forward and declines before reaching the venter. The flanks are otherwise smooth, but the venter is distinctly crenulated. The lectotype differs in no significant respects from material from the Prairie Bluff Chalk. The combination of binodose flanks and a fastigiate venter distinguishes E. labyrinthicus from all others referred to the genus. Eubaculites vagina (Forbes, 1846) (see revision in Kennedy and Henderson, 1992b) has binodose flanks, but is distinguished by a tabulate venter in all but the earliest ontogenetic stages. Other species are discussed by Kennedy (1986d), Kennedy and Henderson (1992b), and Henderson et al. (1992). OCCURRENCE.- Maastrichtian, Prairie Bluff Chalk at localities 15, 40, 46, and 47; Navesink Formation in New Jersey. The types of E. ootacodensis are from Ootacod, South India, and the species also occurs in Madagascar, Zululand (South Africa) and, possibly, Argentina.
Systematics:

46
 Ordo Ammonoidea
  Subordo Ancyloceratina
   Superfamilia Turrilitaceae
    Familia Baculitidae
     Genus Eubaculites
      Species Eubaculites labyrinthicus
Synonym list:
Cobban & Kennedy (1995):
1834 Baculites labyrinthicus Morton. - Morton : p.44 pl. 13; fig. 10
1861 Baculites labyrinthicus Morton. - Gabb : p.394 pl. 3; fig. 1
1866 Baculites vagina var. Ootacodensis Stoliczka. - Stoliczka : p.199 pl. 90; fig. 14 non 15
1895 Baculites vagina var. Ootacodensis Stoliczka. - Kossmat : p.157 pl. 19(5), ?fig. 15 [non fig. 16 = Eubaculites carinatus (Morton, 1834)]
1922 Baculites vagina Forbes. - Cottreau : p.180 pl. 17; fig. 11
1923 Baculites vagina Forbes. - Spengler : p.54 pl. 4; fig. 9
p 1925 Baculites vagina var. Ootacodensis Stoliczka. - Diener : p.63
non 1940 Eubaculites ootacodensis Stoliczka. - Spath : p.49 pl. 1, fig. 3; text-fig. 1b [=E. carinatus]
non 1957 Eubaculites ootacodensis Stoliczka. - Wright : L218 text-fig. 245, 6a-b [=E. carinatus]
non 1959 Eubaculites ootacodensis Stoliczka. - Matsumoto : p.166 pl. 43, fig. 6; pl. 44, fig. 1-3; text-fig. 84-85 [=E. carinatus]
non 1966 Eubaculites ootacodensis Stoliczka. - BRUNNSCHWEILER : p.27 pl. 1; fig. 9-14 [=E. carinatus]
? 1967 Eubaculites sp. . - Leanza : p.55 pl. 1; fig. 6, 7
non 1971 Eubaculites ootacodensis Stoliczka. - COLLIGNON : p.18 pl. 645; fig. 2395 [?=E. carinatus]
1971 Baculites simplex Kossmat. - COLLIGNON : p.15 pl. 645; fig. 2395
non 1974 Eubaculites ootacodensis Stoliczka. - Riccardi : p.388 pl. 1, fig. 1-7; text-fig. 2 [=E. carinatus]
p 1976 Eubaculites ootacodensis Stoliczka. - KLINGER : p.90 pl. 39; fig. 3 only [non: pl. 39, fig. 1; pl. 41, fig. 1, 2; pl. 42, fig. 3, 8; text-fig. 11c (=E. carinatus)]
p 1986 Eubaculites ootacodensis Stoliczka. - Kennedy : p.194
1992 Eubaculites labyrinthicus Morton. - Kennedy & Henderson : p.714 text-fig. 5d-k
1993 Eubaculites labyrinthicus Morton. - KLINGER & Kennedy : p.194 fig. 1-6, 7f-h, 36a-d, 52a-c
1995 Eubaculites labyrinthicus Morton. - Cobban & Kennedy : p. 21, 22, 25 fig. 14.2, 15.2, 15.3, 17.36-17.51
Stratigraphy - relative ages:
Maastrichtian: Cobban & Kennedy (1995)
References:

Morton,S.G.. (1834):
Synopsis of the Organic Remains of the Cretaceous Group of the United States.

Gabb,W.M.. (1861):
Synopsis of the Mollusca of the Cretaceous Formation, including the geographical and stratigraphical range and synonymy . Proceedings of the American Philosophical Society Vol. 8 p. 57-257

Stoliczka,F.. (1866):
The fossil Cephalopoda of the Cretaceous rocks of southern India. Ammonitidae with revision of the Nautilidae etc. . Memoirs of the Geological Survey of India, (l), Palaeontologica Indica10-13 p. 155-216

Kossmat,F.. (1895):
1895-1898. Untersuchungen Ober die Sudindische Kreideformation. . Beiträge zur Paleontologie Österreich-Ungarns und des Orients. p. 9 (1895):97-203

Cottreau,J.. (1922):
Palaeontologie de Madagascar. X.- Fossiles Craetacaes de la Cote Orientale. . Annls. Palaeont Vol. 11 p. 111-192 (1-83)

Spengler,E.. (1923):
Contributions to the palaeontology of Assam . Memoirs of the Geological Survey of India, Palaeontologica IndicaN.S. 8(1) p. 80pp.(4pls.)

Diener,C.. (1925):
Ammonoidea neocretacea . Fossilium Cat, (1: Animalia) Vol. 29 p. 244 pp.

Spath,L.F.. (1940):
On Upper Cretaceous (Maastrichtian) Ammonoidea from Western Australia . Jl. R. Soc. West. Aust. Vol. 26 p. 41-57

Wright,C.W.. (1957):
Family Baculitidae Meek, 1876.
In: Treatise on Invertebrate Paleontology, Part L, Mollusca 4 Eds: Moore, R.C.. p. L218

Matsumoto,T.. (1959):
Upper Cretaceous ammonites of California, Part II. . Memoirs of the Faculty of Science, Kyushu University, Series D, Geology, Special Volume Vol. 1 p. 1-172

BRUNNSCHWEILER,R.o.. (1966):
Upper Cretaceous ammonites from the Carnarvon Basin of Western Australia 1: the heteromorph Lytoceratina. . Bulletin of the Bureau of Mineral Resources, Geology and Geophysics Vol. 58 p. 5-58

Leanza,A.F.. (1967):
Los Baculites de la Provincia de la Pampa con notas acerca de la Edad del Piso Rocanense . Boletin de la Academia Nacional de Ciencias Vol. 46 p. 49-58

COLLIGNON,M.. (1971):
Atlas des fossiles caracteristiques de Madagascar (Ammonites)(Maestrichtian). Vol. 17 p. 82

Riccardi,A.C.. (1974):
Eubaculites Spath (Ammonoidea) del Cretacico Superior de Argentina . Ameghiniana Vol. 11 p. 379-399

KLINGER,H.C.. (1976):
Cretaceous heteromorph ammonites from Zululand . Memoirs of the Geological Survey of the Republic of South Africa Vol. 69 p. 142 pp

Kennedy,W.J.. (1986):
The ammonite fauna of the type Maastrichtian with a revision of Ammonites colligatus Binkhorst, 1861. . Bulletin de I 'Institut Royal des Seiences Naturelles de Belgique, Seiences de la Terre Vol. 56 p. 151-267

Kennedy,W.J.. and Henderson,R.A.. (1992):
Heteromorph ammonites from the Upper Maastrichtian of Pondicherry, south India . Palaeontology Vol. 35 p. 693-731

KLINGER,H.C.. and Kennedy,W.J.. (1993):
retaceous faunas from Zululand and Natal, South Africa. The heteromorph ammonite genus Eubaculites Spath, 1926. . Annals of the South African Museum, 102:185264

Cobban,W.A.. and Kennedy,W.J.. (1995):
Maastrichtian ammonites chiefly from the Prairie Bluff Chalk in Alabama and Mississippi . Journal of Paleontology Vol. 69(5) p. 1-40

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