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Species Favusella washitensis Carsey 1926



Discussion / Comments:
Caron (1985):
3-5 globular chambers form the last whorl. The primary aperture is arched, umbilical- extraumbilical, nearly peripheral, with a narrow lip. The test has a heavily reticulated ornamentation.
Boudagher-Fadel et al. (1997):
Remarks: F. washitensis has commonly four chambers in the last whorl, but occasionally only three and a half chambers are visible on the umbilical side. An intraumbilical aperture occurs with a microperforate wall which has the surface covered by anastomosing ridges, which enclose groups of the microperforations in the cells of the favose reticulation. Pseudomuricae may or may not be present at the confluences of the anastomosing ridges; Plate 4.4, figs 8-9 shows how incipient pseudomuricae are preserved at the intersection of the ridges even in the stratigraphically youngest form of the evolutionary lineage, recalling its conoglobigerine ancestor. The type specimens of F. washitensis, from the Washita group, Texas, were lost from the University of Texas Micropalaeontologica1 Collection and a neotype was proposed by Plummer (1931) from the Del Rio Formation (Early Cenomanian), of the Washita Group, at Shoal Creek, Austin, Texas. Longoria (1974) noted that Carsey's original photograph was a dorsal view, showing four and a half chambers dorsally in the last whorl of the only figured syntype. Consequently, Longoria (1974) believed it was significantly different from the neotype figured by Plummer (1931), as the latter possessed only four chambers visible dorsally in the last whorl. Therefore, Longoria proposed a new neotype from the Grayson Formation of the same locality, which had four and half chambers visible dorsally in the last whorl. The specimens with only four chambers in the last whorl (seen dorsally) were named by Longoria and Gamper (1977) as the new species F. conjksa. It is interesting to note that the specimen figured by Tappan (1940), from the Late Albian Grayson Formation of north Texas, had four chambers visible dorsally in the last whorl, while that figured by Loeblich and Tappan (1961), a topotype from the Cenomanian of the Del Rio Clay, Austin, Texas had four and a half chambers. It is clear from these and other topotypes (e.g. Masters, 1977, plates 25,26, and Michael, 1973, plate 5) that strict topotypes of F. washitensis can have four to four and a half chambers visible dorsally in the last whorl, and three and a half to four chambers visible on the umbilical side. Longoria's proposal (1974) for a second neotype was quite unnecessary, even though it was useful in that a topotypic bullate specimen was imaged. The status of Reticuloglobigerina Reiss (1963) was declared nomen nudum et oblitum by Loeblich and Tappan, 1984, p. 38.
Systematics:

5
 Classis Foraminifera
  Familia Favusellidae
   Genus Favusella
    Species Favusella washitensis

22
  Subordo Globigerinina
   Superfamilia Favusellacea
    Familia Favusellidae
     Genus Favusella
      Species Favusella washitensis
Synonym list:
Banner & Desai (1988):
1926 Globigerina washitensis Carsey. - Carsey :
1988 Favusella washitensis Carsey. - Banner & Desai : 152, 153 pl 2 figs 6-7
Caron (1985):
1926 Globigerina washitensis Carsey. - Carsey : p.44 pl 7 fig 10
1931 Globigerina washitensis Carsey. - Plummer : p.193 pl 13 figs 12a-b (Neotype)
1985 Favusella washitensis Carsey. - Caron : 58,59 figs 25.25-26; 10, 12, 15
Boudagher-Fadel et al. (1997):
1926 Globigerina washitensis Carsey. - Carsey : p.44 pl. 7; fig. 10
1931 Globigerina washitensis Carsey. - Plummer : p.193 pl. 13; fig. 12a, b (first neotype)
1961 Hedbergella washitensis Carsey. - Loeblich & Tappan : pl. 4, fig. 11
1963 Reticuloglobigerina Reiss. - Reiss : p.74
1973 Favusella washitensis Carsey. - Michael : 212, 213 pl. 5; fig. 1-3
1974 Favusella washitensis Carsey. - Longoria : 74, 75 pl. 26; fig. 4-6 (second neotype)
1974 Favusella sp. Longoria. - Longoria : p.74 pl. 26; fig. 8, 9
1977 Favusella confusa Longoria & Gamper. - Longoria & Gamper : 204, 207 pl. 3, fig. 4-6 (paratype), 10-12; pl. 4, fig. 25-27 (holotype)
1977 Globigerina washitensis Carsey. - Masters : p.477 pl. 25, fig. 4; pl. 26, fig. 1-3 (topotype)
1982 Reticuloglobigerina washitensis Carsey. - Banner : p.188 fig. 5.5
1985 Favusella washitensis Carsey. - Caron : p.45 fig. 25.25, 26
1988 Favusella washitensis Carsey. - Banner & Desai : 148-150 pl. 2; fig. 6, 7
1997 Favusella washitensis Carsey. - Boudagher-Fadel et al. : 9, 69, 71 pl. 1.2, fig. 6; pl. 4.3, fig. 7-10; pl. 4.4, fig. 1-9; Figure: 4.1
Was used in synonym list of:
Ascoliella scotiensis Banner & Desai 1988
Ascoliella nitida Michael 1973
References:

Carsey,D.O. (1926):
Foraminifera of the Cretaceous of Central Texas . University of Texas Bulletin Vol. 2612 p. 1-56

Plummer,H.J. (1931):
Some Cretaceous foraminifera in Texas . University of Texas Bulletin Vol. 3101 p. 109-203

Loeblich,A.R. and Tappan,H. (1961):
Cretaceous planktonic foraminifera: Part I-Cenomanian . Micropaleontology Vol. 7 p. 257-304

Reiss,Z.. (1963):
Reclassification of perforate foraminifera . Bulletin of the Geological Survey of Israel Vol. 29 p. 1-28

Michael,F.Y.. (1973):
Planktonic foraminifera from the Comanchean Series (Cretaceous) of Texas . Journal of Foraminiferal Research Vol. 2 p. 200-220

Longoria,J.F. (1974):
Stratigraphic, morphologic and taxonomic studies of Aptian planktonic Foraminifera . Revista Espanola de MicropaleontologíaNo extraord p. 1-107

Masters,B.A. (1977):
Mesozoic Planktonic Foraminifera.
In: Oceanic Micropalaeontology Eds: Ramsay, A.T.S. p. 301-731

Longoria,J.F. and Gamper,M.A. (1977):
Albian planktoinc foraminifera from the Sabinas Basin of northern Mexico . Journal of Foraminiferal Research Vol. 7(3) p. 196-215

Banner,F.T. (1982):
Apects of Micropalaeontology. , A classification and introduction to the Globigerinacea p. 142-239

Caron,M. (1985):
Cretaceous planktic foraminifera.
In: Plankton stratigraphy Eds: Bolli, H.M.Saunders, J.B.Perch-Nielsen, K. p. 17–86

Banner,F.T. and Desai,D. (1988):
A review and revision of the Jurassic-Early Cretaceous Globigerinina, with special reference to the Aptian assemblages of Speeton (North Yorkshire, England) . J. micropalaeontol. Vol. 7(2) p. 143-185

Boudagher-Fadel,M.K.; Banner,F.T.; Whittaker,J.E. and Simmons,M.D. (1997):
Early Evolutionary History of Planktonic Foraminifera. Vol. 1

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