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Species Globorotaloides eovariabilis Huber & Pearson 2006


Diagnosis / Definition:
Pearson et al. (2006):
DESCRIPTION. Type of wall: Normal perforate, coarsely cancellate, succulifer-type wall texture, often with corroded interpore ridges reeking in a remnant wall texture consisting of distinct 'rosettes' around the pores. Test morphology: Test outline lobate, subcircular in axial view, axial periphery rounded to very slightly compressed, biconvex, oval to egg-shaped in edge view; 3-31/2 whorls of slightly inflated chambers arranged in a flattened to slightly elevated trochospire; 14- 15 chambers in adult tests, 41/74 in the final whorl increasing moderately in size; umbilicus shallow to moderately deep and narrow; umbilical suture:-moderately depressed, radial; spiral initially indistinct, later weakly depressed, radial; aperture a low umbilicalextraumbilical arch extending one-third towards peripheral margin, surrounded by a broad lip that extends into the umbilical area. Size: Holotype (USNM 523429) maximum diameter 0.18 mm, breadth 0.10 mm; paratype a (USNM 523430) maximum diameter 0.13 mm, breadth 0.80 mm; paratype b (USNM 523430) maximum diameter 0.15 m. breadth 0.93 mm.
Discussion / Comments:
Pearson et al. (2006):
ETYMOLOGY.- Named as "dawn" vaiabilis because of its superficial resemblance to the much younger species Globorotaloides vuriabilis Bolli. DISTINGUISHING FEATURES.- Differs from Globorotaloides quadrocameratus n. sp. by having 41/2 to 6, rather than 4-41/2, final whorl chambers that are less inflated and increase more gradually in size; differs from Globorotaloides variabilis by its smaller size and less flat inner whorl. DISCUSSION.-This form has been widely recognized at high latitude sites using various species names and open nomenclature. It is included in Globorotaloides because of its coarsely cancellate wall texture and low, sometimes flattened spiral coil. Specimens frequently exhibit corroded interpore ridges with a remnant surface texture consisting of a distinct rosette pattern (PI. 5.4, Figs. 4, 13-15). A high latitude form from the lower Eocene of Site 690 (Maud Rise) that may be related to this species was designated by Stott and Kennett (1990, pl. 5, figs. 3-4) as "Tenuitella" reissi (Loeblich and Tappan). This morphotype has a similar coarsely reticulate wall texture but it is higher spired and the equatorial margin is more compressed. PHYLOGENETIC RELATIONSHIPS.- Descended from Globorotaloides quadrocamerutus n. sp. during the early Eocene. Intermediate morphologies can be found (e.g.,Pl. 5.5, Figs. 13-15). GEOGRAPHIC DISTRIBUTION.-Found at deep-sea sites in southern and northern high latitudes but not at middle or low latitudes. STABLE ISOTOPIC PALEOBIOLOGY.- No data available. TYPE LOCALITY.- The holotype and paratype are from the middle Eocene of ODP Hole 647A (Sample 647A-50R-5,101 cm) in the Labrador Basin. which was assigned to calcareous nannofossil Zone NP 16 by Firth (1989).
Systematics:

35
 Ordo Foraminiferida
  Superfamilia Globigerinaceae
   Familia Globigerinidae
    Genus Turborotalita
     Species Globorotaloides eovariabilis
Synonym list:
Pearson et al. (2006):
1975 Clavigerinella? columbiana Petters. - McKeel & Lipps : p.258 pl. 3, fig. Xa-c (partim; not pl. 4: fig. 6a, b = Parasubbotina eoclava Coxall, Huber, and Pearson) [lower middle Eocene, Elkton Siltstone, Coast Ranges, Oregon]. [Not Petters, 1954.]
? 1979 Globorotalia (Turborotalia) pseudoimitata Blow. - Blow : p. 1104-1105 pl. 101, fig. 1-3 [upper Paleocene Zone P5, DSDP Site 47, Shatsky Rise, northwest Pacific Ocean]; pl. 106, fig. 2-10; pl. 108, fig. 1-3 [lower Eocene Zone P6, Hole 20C, South Atlantic Ocean]
p 1983 Globorotalia munda Jenkins. - Krasheninnikov & Basov : p.841 pl. 10; fig. 101 [lower Eocene, DSDP Site 511, Falkland Plateau, South Atlantic].[Not Jenkins, 1966.]
1990 Globorotaloides aff. testarugosa Jenkins. - Stott & Kennett : p.560 pl. 7; fig. 11 [middle Eocene Zone AE415, ODP Hole 689B, Maud Rise, southern South Atlantic]
1991 Globorotaloides aff. testarugosa Jenkins. - Huber : p.440 pl. 7; fig. 19, 20 [lower Oligocene, Zone AE9, ODP Hole 738B, Kerguelen Plateau, southern Indian Ocean]. [Not Jenkins, 1971 .]
1991 "Planorotalites" reissi Loeblich & Tappan. - Huber : p.440 pl. 4; fig. 18-20 [lower Eocene Zone AE4/5, ODP Hole 738C, Kerguelen Plateau, southern Indian Ocean]. [Not Loeblich and Tappan, 1957.]
1991 Globorotaloides aff. suteri Bolli. - Nocchi et al. : p.268 pl. 4; fig. 4, 5 [lower middle Eocene Zone E8- E9, ODP Site 699, southern South Atlantic]. [Not Bolli, 1957b.]
1992 Globorotaloides suteri Bolli. - Berggren : p.564 pl. 4; fig. 4 [upper Eocene G. suteri Zone ODP Hole 748B, Kerguelen Plateau, southern Indian Ocean]. [Not Bolli, 1957b.]
1992 Globorotaloides sp. 1 Berggren. - Berggren : p.568 pl. 4; fig. 11 [upper Eocene G. suteri Zone, ODP Hole 748B, Kerguelen Plateau, southern Indian Ocean]
2006 Globorotaloides n sp. eovariabilis Huber & Pearson. - Pearson et al. : p.79 pl. 5.4; fig. 1-17
Specimen:
Smithsonian Museum of Natural History, Washington, D.C., Inventory number: USNM 523429
Smithsonian Museum of Natural History, Washington, D.C., Inventory number: USNM 523430a
Smithsonian Museum of Natural History, Washington, D.C., Inventory number: USNM 523430b
References:

McKeel,D.R.. and Lipps,J.H. (1975):
Eocene and Oligocene planktonic foraminifera from the central and southern Oregon coast range . Journal of Foraminiferal Research Vol. 5 p. 1-5

Blow,W.H. (1979):
The Cainozoic Globigerinida. 3 Vols p. 1413 pp

Krasheninnikov,V.A. and Basov,I.A. (1983):
Cenozoic planktonic foraminifers of the Falkland Plateau and Argentine Basin, Deep Sea Drilling Projejt Leg 71.
In: Initial Reports of the Deep Sea Drilling Project Vol. 71 Eds: Ludwig, W.J.Krasheninninov, V.A. p. 821-845

Stott,L.D. and Kennett,J.P. (1990):
Antarctic Paleogene planktonic foraminifer biostratigraphy:ODP Leg 113, Sites 689 and 690.
In: Proceedings of the Ocean Drilling Program Vol. 113

Nocchi,M.; Amici,E. and Premoli Silva,I. (1991):
Planktonic Foraminiferal Biostratigraphy and Paleoenvironmental Interpretation of Paleocene Faunas from the Subantartic Transect, Leg 114.
In: Proceedings of the Ocean Drilling Program, Scientific Results Vol. 114 Eds: Ciesielski, P.F.Kristoffersen, Y.Al, E. p. 233-279

Huber,B.T. (1991):
Paleogene and early Neogene planktonic foraminifer biostratigraphy of Sites 738 and 744, Kerguelen Plateau (southern Indian Ocean).
In: Proceedings of the Ocean Drilling Program, Scientific Results Vol. 119 Eds: Leckie, R.M.Sigurdsson, H.Acton, G.D.Draper, G. p. 427-449

Berggren,W.A. (1992):
Paleogene Planktonic Foraminifera Magnetobiostratigraphy of the Southern Kerguelen Plateau (Sites 747-749).
In: Proceedings of the Ocean Drilling Program, Scientific Results Vol. 120

Pearson,P.N.; Olsson,R.K.; Hemleben,C.; Huber,B.T. and Berggren,W.A. (2006):
Atlas of Eocene Planktonic Foraminifera. p. 1-513
Anonymous: Unedited TaxonConcept data
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