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Species Globigerinatheka euganea Proto Decima & Bolli 1970



Diagnosis / Definition:
Pearson et al. (2006):
DESCRIPTION. Type of wall: Spinose, cancellate, with rather dense pores, about 0.08 mm in diameter. Test morphology: Test almost spherical, mainly 2 to possibly 2 1/2 early whorls coiled in a low trochospire consisting of four to six inflated chambers; chambers initially increase very slowly in size as they are added, then more rapidly, becoming more globular; the last 3 chambers, which increase very rapidly in size, are streptospirally coiled with the last chamber tending to be hemispherical and covering the umbilicus of previous whorls; the last chamber may be of the same size as the penultimate one or larger; sutures distinct, weakly depressed in the earlier whorl becoming clearly depressed in the intermediateflhorls, tending to be less depressed between the last chambers; primary aperture covered in the adult stage; two to three, occasionally four small arched to subcircular secondary apertures along the base of the last chamber at the junctions of previous chambers, that may be covered by small bullae. Size: Maximum diameter of holotype 0.53 mm.
Discussion / Comments:
Pearson et al. (2006):
DISTINGUISHING FEATURES.- Globigerinatheka euganea has a compact, spherical test, with small, numerous secondary apertures in some specimens. It differs from G. curryi by having a more compact test, a tighter inner coil and more numerous and smaller secondary apertures. It differs from G. luterbacheri by having a more spherical test and less depressed sutures; it differs from beckmanni by less numerous, slightly larger secondary apertures, rarely recorded along the inner spire, and by the absence of area1 openings. DISCUSSION.- At first sight G. euganea looks similar to Cushman's original drawing of the G. mexicana holotype, as redrawn by Blow and Saito (1968a). However, the SEM photographs of the mexicana holotype clearly show that the two species are not conspecific. PHYLOGENETIC RELATIONSHIPS.- According to Bolli (1972), G. euganea is the intermediate member of the curryi - euganea - beckmanni lineage. GEOGRAPHIC DISTRIBUTION.- Low to midlatitudes. Large specimens mainly in tropical regions or warmer mid latitudes. STABLE ISOTOPE PALEOBIOL0GY.- No data available.
Systematics:

35
 Ordo Foraminiferida
  Superfamilia Globigerinaceae
   Familia Globigerinidae
    Genus Globigerinatheka
     Species Globigerinatheka euganea
Synonym list:
Pearson et al. (2006):
1927 Globigerina mexicana Cushman. - Cushman : pl. 26; fig. 16, 17 [Mexico]; [Not Cushman, 1925]
p 1953 Globigerinoides mexicana Cushman. - Beckmann : p.395 pl. 25; fig. 15, 16 (not figs. 17-19 = 0. beckmanni); [Eocene, Oceanic Fm., Barbados]; [Not Cushman, 1925]
1960 Globigerapsis mexicana Cushman. - Todd & Low : pl. 259; fig. 7 [drill hole, Eniwetok, central Pacific Ocean]
1962 Globigerapsis mexicana Cushman. - Saito : pl. 34; fig. 7a-c (not 6a-b) [Eocene, Hahajima, Hillsborough Island, Japan]; [Not Cushman, 1925]
1970 Globigerinatheka euganea Proto Decima & Bolli. - Proto Decima & Bolli : p.894 pl. I, fig. 6-11; pl. III, fig. 3-5; text-fig. 8-15, 31-31a, 43-45b (holotype); pl. I, fig. 7 (paratype); pl. I, fig. 8; pl. III, fig. 4; text-fig. 40-42a [middle Eocene Orbulinoides beckmanni Zone, Euganei marls, Cava Zillo, northern Italy]
1970 Globigerapsis cf. kugleri Bolli et al.. - Baumann : pl. 1; fig. 13a, b [middle Eocene Glohorotalia centralis Zone, probably P12, Bottaccione Section, Gubbio, central Italy]; [Not Bolli, Loeblich and Tappan, 1957]
1970 Globigerinatheka barri Brönnimann. - Baumann : pl. 3; fig. 1 [middle Eocene Globigerapsis mexicana Zone, Scalette Section, Umbria, central Italy]; [Not Bronnimann, 1952]
1972 Globigerinatheka subconglobata euganea Proto Decima & Bolli. - Bolli : p.132 text-fig. 31-34 (redrawn from Proto Decima and Bolli, 1970); text-fig. 35 (redrawn from Cushman, 1927); text-fig. 36-38 (redrawn from Beckmann, 1953); text-fig. 39 (redrawn from Todd and Low, 1960); text-fig. 40-42 (redrawn from Saito, 1962)
1975 Globigerinatheka euganea Proto Decima & Bolli. - Toumarkine : pl. 4; fig. 13, 14 [middle Eocene Orbulinoides beckmanni Zone, DSDP Site 3 13, Mid-Pacific Mountains]
1978 Globigerinatheka subconglobata euganea Proto Decima & Bolli. - Toumarkine : pl. 3; fig. 6-7 [middle Eocene Orbulinoides beckmanni Zone, DSDP Site 363, South Atlantic Ocean]
p 1979 Globigerapsis kugleri euganea Decima & Bolli. - Blow : p.1149 pl. 181, fig. 6; pl. 183, fig. 5, 6; pl. 185, ?fig. 6 [middle Eocene Zone P1l, DSDP Hole 21A, South Atlantic Ocean]; pl. 186 (partim), ?fig. 4 (not fig. 5 = G. euganea), [middle Eocene Zone P12, DSDP Site 19, South Atlantic Ocean]; pl. 192 (partim), fig. 4 (not fig. 2, 3), [middle Eocene Zone P13, Kilwa area, Tanzania]
1979 Globigerapsis beckmanni Saito. - Blow : p. 1137-1138 pl. 192; fig. 6-9, ?fig. 5 [middle Eocene Zone P13, Kilwa area, RS.3 11, Tanzania]; [Not Saito, 1962]
1983 Globigerinatheka subconglobata euganea Proto Decima & Bolli. - Toumarkine : pl. 21; fig. 8-10 [fig. 8, middle Eocene Orbulinoides beckmanni Zone, DSDP Site 313, Mid Pacific Mountains; fig. 9, middle Eocene Morozovella lehrzeri zone, DSDP Site 313, Mid Pacific Mountains; fig. 10, middle Eocene Orbulinoides beckmanni Zone, DSDP Site 363, S
? 1983 Globigerinatheka barri Brönnimann. - Pujol : pl. 8; fig. 12 [upper Eocene Zone P15-16, DSDP Hole 516F. Rio Grande Rise, South Atlantic Ocean]. [Not Bronnimann, 1952]
2006 Globigerinatheka euganea Proto Decima & Bolli. - Pearson et al. : p.182 pl. 7.4; fig. 1-8, 11, 12 (Pl. 7.4, Figs. l, 2: reillustration of holotype and paratype of Globigerinatheka euganea Proto Decima and Bolli)
Specimen:
National Museum of Natural History, Basel, Inventory number: C 26543
National Museum of Natural History, Basel, Inventory number: C 26529
National Museum of Natural History, Basel, Inventory number: C 26530
National Museum of Natural History, Basel, Inventory number: C 26531
National Museum of Natural History, Basel, Inventory number: C 26544
References:

Cushman,J.A. (1927):
Some characteristic Mexican fossil foraminifera . Journal of Paleontology Vol. 1 p. 147-172

Beckmann,J.P. (1953):
Die Foraminiferen der Oceanic Formation (Eocaen-Oligocaen) von Barbados. KI. Antillen . Eclogae Geologicae Helvetiae Vol. 46 p. 301-412

Todd,R. and Low,D. (1960):
Smaller foraminifera from Eniwetok drill hole . U.S. Geological Survey, Professional paper 260-X p. 799-861

Saito,T. (1962):
Eocene planktonic foraminifera from Hahajima (Hillsborough Island) . Trans. Proc. Paleontol. Soc. Japan, news series Vol. 45 p. 209-225

Baumann,P.. (1970):
Mikropalaentologische und stratigraphische Untersuchungen der obereozaenen-oligozaenen Scaglia im zentralen Apennin (Italien) . Eclogae Geologicae Helvetiae Vol. 63 p. 1133-1211

Proto Decima,F. and Bolli,H.M. (1970):
Evolution and variability of Orbulinoides beckmanni (Saito) . Ecoglae geologicae Helvetiae Vol. 63 p. 883-905

Bolli,H.M. (1972):
The Genus Globigerinatheka Brönnimann . Journal of Foraminiferal Research Vol. 2 p. 109-136

Toumarkine,M. (1975):
Middle and Late Eocene planktonic foraminifera from the northwestern Pacific, Leg 32 of the Deep Sea Drilling Project.
In: Initial Reports of the Deep Sea Drilling Project Vol. 32 Eds: Larson, R.L.Moberly, R. p. 735-751

Toumarkine,M. (1978):
Planktonic Foraminiferal Biostratigraphy of the Paleogene of Sites 360 to 364 and the Neogene of Sites 362A, 363, and 364 Leg 40.
In: Initial Results of the Deep Sea Drilling Project Vol. 40

Blow,W.H. (1979):
The Cainozoic Globigerinida. 3 Vols p. 1413 pp

Toumarkine,M. (1983):
Les Foraminiferes planctoniques de l'Eocéne moyen et supérieur des regions tropicales ŕ temperées chaudes.
In: Thése de Doctorat d'Etat čs Sciences Vol. 6(83-05) p. 219

Pujol,C. (1983):
Cenozoic Planktonic Foraminiferal Biostratigraphy of the Southwestern Atlantic (Rio Grande Rise): Deep Sea Drilling Project Leg 72.
In: Initial Reports of the Deep Sea Drilling Project Vol. 72 Eds: Barker, P.F.Johnson, D.A. p. 623-673

Pearson,P.N.; Olsson,R.K.; Hemleben,C.; Huber,B.T. and Berggren,W.A. (2006):
Atlas of Eocene Planktonic Foraminifera. p. 1-513

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