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Species Acarinina coalingensis Cushman & Hanna 1927


Discussion / Comments:
Van Eijden & Smit (1991):
Remarks. This species is the central form in a plexus of four-chambered forms with more or less appressed chambers and a rounded or somewhat angular periphery (A. soldadoensis angulosa (Bolli, 1957b), A. esnaensis LeRoy, 1953, A. nitida (Martin, 1943), A. pseudotopilensis (Subbotina, 1953), and A. soldadoensis (Brönnimann, 1952c)). Because these are connected by intermediate forms and most have similar ranges, we counted them together. A. primitiva (Finlay, 1947) (= Globoquadrina primitiva) is a junior synonym.
Pearson et al. (2006):
See Olsson and others (1999, p. 47) for synonymy list and taxonomic discussion (and also discussion, below). DISCUSSION.- Berggren and Norris (1997) and Olsson and others (l999) discussed the gradual evolution of the coalingensis-primitiva lineage and included primitiva in synonymy with coalingensis. Having studied populations of A. primitiva in the type Hampden Beach Formation, New Zealand, we now conclude that they are not synonymous; see also under A. pyimitiva. We restrict the concept of coalingensis to the robust, triangular-subquadrate tests with (predominantly) globular chambers (typified by Globigerina coalingensis Cushman and Hanna) and its junior synonym Acarinina triplex Subbotina. STRATIGRAPHIC RANGE.- Zone P4c-E7; (not to P14, = E13 of this study, as given by Olsson and others, 1 999).
Systematics:

1
 Superregnum Eukaryota
  Regnum Protoctista
   Phylum Ciliophora
    Subphylum Postciliodesmatophora
     Ordo Globigerinida
      Superfamilia Globorotaliaceae
       Superfamilia Nonionacea
        Familia Truncorotaloididae
         Genus Acarinina
          Species Acarinina coalingensis

17
 Classis Foraminifera
  Familia Truncorotaloididae
   Genus Acarinina
    Species Acarinina coalingensis

19
  Ordo Foraminiferida
   Superfamilia Globorotaliaceae
    Familia Truncorotaloididae
     Genus Acarinina
      Species Acarinina coalingensis
       Species Morozovella conicotruncata

32
 Ordo Foraminiferida
  Familia Globorotaliidae
   Genus Acarinina
    Species Acarinina coalingensis

35
  Ordo Foraminiferida
   Superfamilia Globigerinaceae
    Familia Truncorotaloididae
     Genus Acarinina
      Species Acarinina coalingensis
Synonym list:
Berggren & Norris (1997):
1927 Globigerina coalingensis Cushman & Hanna. - Cushman & Hanna : p. 219 pl. 14, fig. 4 (lower Eocene, California)
1947 Globoquadrina primitiva Finlay. - Finlay : p. 291 pl. 8, figs. 129-134.
1952 Globigerina primitiva Finlay. - Brönnimann : p. 11 pl. 1, figs. 10-12 (Trinidad; Soldado and Lizard Springs Formations).
1953 Globoquadrina primitiva Finlay. - Hornibrook : p. 437.
1953 Acarinina triplex Subbotina. - Subbotina : p. 230 pl. 23, figs. 1-5 (holotype from Gl. marginodentata Subzone of Gl. subbotinae Zone, lower Eocene, Khieu River, Nal'chik, N Caucasus).
1957 Globigerina primitiva Finlay. - Bolli : p. 71 pl. 15 figs. 6-8
1957 Globigerina cf. soldadoensis Brönnimann. - Loeblich & Tappan (non Brönnimann, 1952) : p. 182 pl. 53, fig. 1 (Zone P4; Vincentown Fm., NJ, upper Paleocene)
1957 Globigerina inaequispira Subbotina. - Loeblich & Tappan (non Subbotina, 1953) : p. 181 pl. 61, fig. 3 (Zone P4, Nanafalia Fm., Alabama, upper Paleocene).
1959 Globigerina coalingensis Cushman & Hanna. - Mallory : p. 249 pl. 34, figs. 6 a-c (Gredal, Devil's Den, Ulatisian Stage [Ypresian], California).
1961 Globigerina primitiva Finlay. - Hornibrook : p. 148.
1962 Globorotalia (Acarinina) primitiva Finlay. - von Hillebrandt : p. 141 pl. 14, figs. 2,4 (Zone G, lower Eocene, Austria).
1965 Globoquadrina primitiva Finlay. - Jenkins : p. 269 pl. 1, fig. (outline drawing of Finlay's holotype).
1965 Pseudogloboquadrina primitiva Finlay. - Jenkins : p. 1124-1125 fig. 9 (nos. 81-83 (holotype) and 84-86; Bortonian, middle Eocene, New Zeland).
1970 Acarinina coalingensis Cushman & Hanna. - Berggren : p. 152 pl. 1, figs. 27-29 (Zone NP 11/12, stratigraphically equivalent to Zone P6/7; Lower Eocene 3, Katharinenhof, Fehmarn, N Germany).
1971 Globigerina primitiva Finlay. - Postuma : p. 154 7 figured specimens (Hampden, Otago Province, South Island, New Zealand, lower Eocene).
1971 Pseudogloboquadrina primitiva Finlay. - Jenkins : p. 170 pl. 18, figs. 551-561 (original types refigured, 550-560).
1975 Globigerina primitiva Finlay. - Stainforth et al. : p. 215 Fig. 75 (1-3) (Gl. formosa Zone, Lodo Formation, California).
1977 Acarinina coalingensis Cushman & Hanna. - Berggren : p. 251 pl. 10
1979 Acarinina triplex Subbotina. - Blow : pl. 1, figs. 11-12 (Zone Pll, middle Eocene, DSDP Hole 523, S Atlantic).
1979 Globorotalia (Acarinina) primitiva Finlay. - Blow : p. 949 pl. 143, figs. 6-9 (Zone P8, lower Eocene, DSDP Hole 20C); pl. 249, figs. 1-4 (lower Bortonian Stage, middle Eocene, New Zealand).
1979 Globorotalia (Acarinina) triplex Subbotina. - Blow : p. 963 pl. 97, figs. 8,9 (Zone P5, Lindi area, Tanzania, E Africa).
1983 Pseudogloboquadrina primitiva Finlay. - Krasheninnikov & Basov : p. 840 pl. 9, figs. 4-7 (middle Eocene, DSDP Hole 512, Ewing Bank, S Atlantic).
1990 Acarinina primitiva Finlay. - Stott & Kennett : p. 559 pl. 6, figs. 11,12 (Zone AP8, lower Eocene, ODP Hole 689B, Maude Rise, Weddell Sea, Antarctic Ocean).
1991 Acarinina coalingensis Cushman & Hanna. - Huber : pl. 3, fig. 2 (Zone AP6b/AP7; lower Eocene, ODP Site 738, Kerguelen Plateau, S Indian Ocean).
1991 Acarinina primitiva Finlay. - Huber : p. 439 pl. 3, fig. l (Zone AP10, middle Eocene, ODP Site 738, Kerguelen Plateau, S Indian Ocean).
1992 Acarinina coalingensis Cushman & Hanna. - Berggren : p.563 pl. 2, fig. 3 (Ac. wilcoxensis Zone, lower Eocene, ODP Site 749, Kerguelen Plateau, S Indian Ocean).
1992 Acarinina primitiva Finlay. - Berggren : p. 563 pl. 2, figs. 4,5 (Zone P6-7, ODP Site 748, Kerguelen Plateau, S Indian Ocean).
1993 Acarinina triplex Subbotina. - Lu & Keller : pl. 2, figs. 4.5 (Zone AP 6A, ODP 119/738C/9R/1: 15-17cm; lowermost Eocene; S Indian Ocean; Kerguelen Plateau).
1993 Acarinina primitiva Finlay. - Pearson et al. : pl. 1, fig. 19 (Zone Pll, middle Eocene, DSDP Hole 523, S Adantic).
1993 Acarinina primitiva Finlay. - Lu & Keller : pl 3, figs. 1,2 (Zone AP7, ODP 119/738C/5R/2: 57-59cm; lower Eocene; S Indian Ocean (Kerguelen Plateau).
1995 Acarinina triplex Subbotina. - Basov : p. 165 pl. 1, figs. 16, 17 (M. aragonensis s.l. Zone; ODP 145/883B/82X/5: 120-122cms; Detroit Seamount, NW Pacific).
1995 Acarinina primitiva Finlay. - Basov : p. 165 pl. 1, figs. 11-13 (ODP 145/883B/86X/4: 120-122cms; Detroit Seamount, NW Pacific; NB: not recorded at this level in Table 1, p. 160).
1997 Acarinina coalingensis Cushman & Hanna. - Berggren & Norris : p. 95 Plate 12 Figure 16-20, 22, 23
Olsson et al. (1999):
1927 Globigerina coalingensis Cushman & Hanna. - Cushman & Hanna : p. 219 pl. 14, fig. 4 (lower Eocene, California).-Mallory, 1959, p. 46, pl. 34, fig. 6 (lower Eocene, California)
1947 Globoquadrina primitiva Finlay. - Finlay : p. 291 pl. 8, figs. 129-134
1952 Globigerina primitiva Finlay. - Brönnimann : p. 11 pl. 1, figs. 10-12 (Trinidad; Soldado and Lizard Springs Fms.)
1953 Acarinina triplex Subbotina. - Subbotina : p. 230 pl. 23, figs. 1-5 (holotype from G. marginodentata Subzone of G. Subbotinae Zone, lower Eocene, Khieu River, Nal'chik, N Caucasus)
1953 Globoquadrina primitiva Finlay. - Hornibrook : p. 437.
1957 Globigerina primitiva Finlay. - Bolli : p. 71 pl. 15, figs. 6-8 (G. rex Zone, Trinidad)
1957 Globigerina inaequispira Subbotina. - Loeblich & Tappan (non Subbotina, 1953) : p. 181 pl. 61, fig. 3 (Zone P4, Nanafalia Fm., Alabama, upper Paleocene)
1962 Globorotalia (Acarinina) primitiva Finlay. - von Hillebrandt : p. 141 pl. 14, figs. 2,4 (Zone G, lower Eocene, Austria)
1965 Globoquadrina primitiva Finlay. - Jenkins : p. 269 pl. 1, fig. (outline drawing of Finlay's holotype)
1966 Pseudogloboquadrina primitiva Finlay. - Jenkins : p. 1124-1125 fig. 9 [nos. 81-83 (holotype) and 84-86; Bortonian, middle Eocene, New Zealand]
1969 Acarinina coalingensis Cushman & Hanna. - Berggren : p. 152 pl. 1, figs. 27-29 (Zone NP 11/12, stratigraphically equivalent to Zone P6/7; Lower Eocene 3, Katharinenhof, Fehmarn, N Germany)
1979 Acarinina triplex Subbotina. - Blow : p. 963 pl.97, figs. 8, 9 (Zone P5, Linde area, Tanzania, E Africa)
1979 Globorotalia (Acarinina) primitiva Finlay. - Blow : p. 949 pl. 143, figs. 6-9 Zone P8, lower Eocene, DSDP Hole 20C/5/4: 77-79 cm); pl. 249, figs. 1-4 (lower Bortonian, middle Eocene, New Zealand)
1979 Globorotalia (Acarinina) triplex Subbotina. - Blow : p. 963 pl. 97, figs. 8,9 (Zone P5, Lindi area, Tanzania, E Africa).
1990 Acarinina primitiva Finlay. - Stott & Kennett : p. 559 pl. 6, figs. 11,12 (Zone AP8, lower Eocene, ODP Hole 689B/21H/1: 110-114 cm, Maud Rise, Weddell Sea, Antaractic Ocean)
1991 Acarinina coalingensis Cushman & Hanna. - Huber : p. 439 pl. 3, fig. 2 (Zone AP6b/AP7; lower Eocene, ODP Hole 738C/6R: 247.83 msbf, Kerguelen Plateau, Southern Indian Ocean)
1991 Acarinina primitiva Finlay. - Huber : p. 439 pl. 3 fig. 1 ( Zone AP10, middle Eocene, ODP Site 738B/15X: 126.70 msbf, Kerguelen Plateau, Southern Indian Ocean)
1992 Acarinina coalingensis Cushman & Hanna. - Berggren : p.563 pl. 2, fig. 3 (A. wilcoxensis Zone, lower Eocene, ODP Site 749C/10R/1: 40-44 cm, Kerguelen Plateau, Southern Indian Ocean)
1992 Acarinina primitiva Finlay. - Berggren : p. 563 pl. 2, figs. 4,5 (Zone P6-7, ODP Hole 748B/18H/3: 80-84 cm, Kerguelen Plateau, Southern Indian Ocean)
1993 Acarinina triplex Subbotina. - Pearson et al. : pl. 1, figs. 11-12 (Zone P11, middle Eocene, DSDP Hole 523, South Atlantic Ocean)
1993 Acarinina primitiva Finlay. - Pearson et al. : pl. 1 fig. 19 (Zone P11, middle Eocene, DSDP Hole 523, South Atlantic Ocean)
1993 Acarinina primitiva Finlay. - Lu & Keller : pl 3, figs. 1, 2 ( Zone AP7, ODP Hole 738C/5R/2: 57-59 cm, lower Eocene; Kerguelen Plateau, Southern Indian Ocean)
1995 Acarinina triplex Subbotina. - Lu & Keller : pl. 2, figs. 4, 5 (Zone AP 6A, ODP Hole 738C/9R/1: 15-17 cm; lowermost Eocene; Southern Indian Ocean (Kerguelen Plateau)
Van Eijden & Smit (1991):
1927 Globigerina coalingensis Cushman & Hanna. - Cushman & Hanna : p.219 pl. 14, figs. 4a-b
1991 Acarinina coalingensis Cushman & Hanna. - Van Eijden & Smit : p.121 pl. 3, figs. 14.20; pl. 4, fig. 6
Pearson et al. (2006):
2006 Acarinina coalingensis Cushman & Hanna. - Pearson et al. : p.277 pl. 9.7; fig. 1-16 (Pl. 9.7, Figs. 1-3: new SEMs of holotype of Globigerina coalingensis Cushman and Hanna)
Stratigraphy - absolute ages:
FAD: 56.5 ± 0 [Ma], Berggren et al. (1995) DSDP Leg 43 Site 384
References:

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Foraminifera from the Eocene near San Diego, California . San Diego Natural History Society, Transactions Vol. 5 p. 45-64

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The genera Globigerina and Globorotalia in the Paleocene-Lower Eocene Lizard Springs Formation of Trinidad, B.W.I . Bulletin of the U.S. National Museum Vol. 215 p. 61-82

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Planktonic Foraminifera of Paleocene and Early Eocene Age from the Gulf and Atlantic Coastral Plains.
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Anonymous: Unedited TaxonConcept data
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